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The Genetic CodesCompiled by Andrzej (Anjay) Elzanowski and Jim Ostell NCBI takes great care to ensure that the translation for each coding sequence (CDS) present in GenBank records is correct. Central to this effort is careful checking on the taxonomy of each record and assignment of the correct genetic code (shown as a /transl_table qualifier on the CDS in the flat files) for each organism and record. This page summarizes and references this work. The synopsis presented below is based primarily on the reviews by Osawa et al. (1992) and Jukes and Osawa (1993). Listed in square brackets [] (under Systematic Range) are tentative assignments of a particular code based on sequence homology and/or phylogenetic relationships. The print-form ASN.1 version of this document, which includes all the genetic codes outlined below, is also available here. Detailed information on codon usage can be found at the Codon Usage Database. GenBank format by historical convention displays mRNA sequences using the DNA alphabet. Thus, for the convenience of people reading GenBank records, the genetic code tables shown here use T instead of U. The following genetic codes are described here:
1. The Standard Code (transl_table=1)By default all transl_table in GenBank flatfiles are equal to id 1, and this is not shown. When transl_table is not equal to id 1, it is shown as a qualifier on the CDS feature.
AAs = FFLLSSSSYY**CC*WLLLLPPPPHHQQRRRRIIIMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = ---M---------------M---------------M---------------------------- Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Initiation Codon:AUG
Alternative Initiation Codons 2. The Vertebrate Mitochondrial Code (transl_table=2)
AAs = FFLLSSSSYY**CCWWLLLLPPPPHHQQRRRRIIMMTTTTNNKKSS**VVVVAAAADDEEGGGG Starts = --------------------------------MMMM---------------M------------ Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 2 Standard AGA Ter * Arg R AGG Ter * Arg R AUA Met M Ile I UGA Trp W Ter * Alternative Initiation Codon:Bos: AUAHomo: AUA, AUU Mus: AUA, AUU, AUC Coturnix, Gallus: also GUG (Desjardins and Morais, 1991) Systematic Range:Vertebrata
Comment:
3. The Yeast Mitochondrial Code (transl_table=3)
AAs = FFLLSSSSYY**CCWWTTTTPPPPHHQQRRRRIIMMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = ----------------------------------MM---------------------------- Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 3 Standard AUA Met M Ile I CUU Thr T Leu L CUC Thr T Leu L CUA Thr T Leu L CUG Thr T Leu L UGA Trp W Ter * CGA absent Arg R CGC absent Arg R Systematic Range:Saccharomyces cerevisiae, Candida glabrata, Hansenula saturnus, and Kluyveromyces thermotolerans (Clark-Walker and Weiller, 1994)
Comments:
4. The Mold, Protozoan, and Coelenterate Mitochondrial Code and the Mycoplasma/Spiroplasma Code (transl_table=4)
AAs = FFLLSSSSYY**CCWWLLLLPPPPHHQQRRRRIIIMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = --MM---------------M------------MMMM---------------M------------ Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 4 Standard UGA Trp W Ter * Alternative Initiation Codons:Trypanosoma: UUA, UUG, CUGLeishmania: AUU, AUA Tertrahymena: AUU, AUA, AUG Paramecium: AUU, AUA, AUG, AUC, GUG, GUA(?) (Pritchard et al., 1990) Systematic Range:Bacteria: The code is used in Entomoplasmatales and Mycoplasmatales (Bove et al. 1989). The situation in the Acholeplasmatales is unclear. Based on a study of ribosomal protein genes, it had been concluded that UGA does not code for tryptophan in plant-pathogenic mycoplasma-like organisms (MLO) and the Acholeplasmataceae (Lim and Sears, 1992) and there seems to be only a single tRNA-CCA for tryptophan in Acholeplasma laidlawii (Tanaka et al. 1989). In contrast, in a study of codon usage in Phytoplasmas, it was found that 30 out of 78 ORFs analyzed translated better with code 4 (UGA for tryptophan) than with code 11 while the remainder showed no differences between the two codes (Melamed et al. 2003). In addition, the coding reassignment of UGA Stop --> Trp can be found in an alpha-proteobacterial symbiont of cicadas: Candidatus Hodgkinia cicadicola (McCutcheon et al. 2009).Fungi: Emericella nidulans, Neurospora crassa, Podospora anserina, Acremonium (Fox, 1987), Candida parapsilosis (Guelin et al., 1991), Trichophyton rubrum (de Bievre and Dujon, 1992), Dekkera/Brettanomyces, Eeniella (Hoeben et al., 1993), and probably Ascobolus immersus, Aspergillus amstelodami, Claviceps purpurea, and Cochliobolus heterostrophus. Other Eukaryotes: Gigartinales among the red algae (Boyen et al. 1994), and the protozoa Trypanosoma brucei, Leishmania tarentolae, Paramecium tetraurelia, Tetrahymena pyriformis and probably Plasmodium gallinaceum (Aldritt et al., 1989). Metazoa: Coelenterata (Ctenophora and Cnidaria) Comments:
5. The Invertebrate Mitochondrial Code (transl_table=5)
AAs = FFLLSSSSYY**CCWWLLLLPPPPHHQQRRRRIIMMTTTTNNKKSSSSVVVVAAAADDEEGGGG Starts = ---M----------------------------MMMM---------------M------------ Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Comment:The codon AGG is absent in Drosophila.Differences from the Standard Code:Code 5 Standard AGA Ser S Arg R AGG Ser S Arg R AUA Met M Ile I UGA Trp W Ter * Alternative Initiation Codons:AUA, AUUAUC: Apis (Crozier and Crozier, 1993) GUG: Polyplacophora (Boore and Brown, 1994 GenBank Accession Number: U09810) UUG: Ascaris, Caenorhabditis Systematic Range:Nematoda: Ascaris, Caenorhabditis;Mollusca: Bivalvia (Hoffmann et al., 1992); Polyplacophora (Boore and Brown, 1994) Arthropoda/Crustacea: Artemia (Batuecas et al., 1988); Arthropoda/Insecta: Drosophila [Locusta migratoria (migratory locust), Apis mellifera (honeybee)] Comments:
6. The Ciliate, Dasycladacean and Hexamita Nuclear Code (transl_table=6)
AAs = FFLLSSSSYYQQCC*WLLLLPPPPHHQQRRRRIIIMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = -----------------------------------M---------------------------- Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 6 Standard UAA Gln Q Ter * UAG Gln Q Ter * Systematic Range:Ciliata: Oxytricha and Stylonychia (Hoffman et al. 1995), Paramecium, Tetrahymena, Oxytrichidae and probably Glaucoma chattoni.Dasycladaceae: Acetabularia (Schneider et al., 1989) and Batophora (Schneider and de Groot, 1991).
Diplomonadida: Comment:
Tables 7 and 8 have been deleted9. The Echinoderm and Flatworm Mitochondrial Code (transl_table=9)
AAs = FFLLSSSSYY**CCWWLLLLPPPPHHQQRRRRIIIMTTTTNNNKSSSSVVVVAAAADDEEGGGG Starts = -----------------------------------M---------------M------------ Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 9 Standard AAA Asn N Lys K AGA Ser S Arg R AGG Ser S Arg R UGA Trp W Ter * Systematic Range:Asterozoa (starfishes) (Himeno et al., 1987) Echinozoa (sea urchins) (Jacobs et al., 1988; Cantatore et al., 1989) Rhabditophora among the Platyhelminthes (Telford et al. 2000)10. The Euplotid Nuclear Code (transl_table=10)
AAs = FFLLSSSSYY**CCCWLLLLPPPPHHQQRRRRIIIMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = -----------------------------------M---------------------------- Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 10 Standard UGA Cys C Ter * Systematic Range:Ciliata: Euplotidae (Hoffman et al. 1995).11. The Bacterial, Archaeal and Plant Plastid Code (transl_table=11)
AAs = FFLLSSSSYY**CC*WLLLLPPPPHHQQRRRRIIIMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = ---M---------------M------------MMMM---------------M------------ Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Systematic Range and Comments:Table 11 is used for Bacteria, Archaea, prokaryotic viruses and chloroplast proteins. As in the standard code, initiation is most efficient at AUG. In addition, GUG and UUG starts are documented in Archaea and Bacteria (Kozak 1983, Fotheringham et al. 1986, Golderer et al. 1995, Nolling et al. 1995, Sazuka & Ohara 1996, Genser et al. 1998, Wang et al. 2003). In E. coli, UUG is estimated to serve as initiator for about 3% of the bacterium's proteins (Blattner et al. 1997). CUG is known to function as an initiator for one plasmid-encoded protein (RepA) in Escherichia coli (Spiers and Bergquist, 1992). In addition to the NUG initiations, in rare cases Bacteria can initiate translation from an AUU codon as e.g. in the case of poly(A) polymerase PcnB and the InfC gene that codes for translation initiation factor IF3 (Polard et al. 1991, Liveris et al. 1993, Sazuka & Ohara 1996, Binns & Masters 2002). The internal assignments are the same as in the standard code though UGA codes at low efficiency for Trp in Bacillus subtilis and, presumably, in Escherichia coli (Hatfiled and Diamond, 1993). 12. The Alternative Yeast Nuclear Code (transl_table=12)
AAs = FFLLSSSSYY**CC*WLLLSPPPPHHQQRRRRIIIMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = -------------------M---------------M---------------------------- Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 12 Standard CUG Ser Leu Alternative Initiation Codons:CAG may be used in Candida albicans (Santos et al., 1993).Systematic Range:Endomycetales (yeasts): Candida albicans, Candida cylindracea, Candida melibiosica, Candida parapsilosis, and Candida rugosa (Ohama et al., 1993).Comment:
13. The Ascidian Mitochondrial Code (transl_table=13)
AAs = FFLLSSSSYY**CCWWLLLLPPPPHHQQRRRRIIMMTTTTNNKKSSGGVVVVAAAADDEEGGGG Starts = ---M------------------------------MM---------------M------------ Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 13 Standard AGA Gly G Arg R AGG Gly G Arg R AUA Met M Ile I UGA Trp W Ter * Systematic range and Comments:There is evidence from a phylogenetically diverse sample of tunicates (Urochordata) that AGA and AGG code for glycine. In other organisms, AGA/AGG code for either arginine or serine and in vertebrate mitochondria they code a STOP. Evidence for glycine translation of AGA/AGG has been found in Pyura stolonifera (Durrheim et al. 1993), Halocynthia roretzi (Kondow et al. 1999, Yokobori et al., 1993, Yokobori et al. 1999) and Ciona savignyi (Yokobori et al. 2003). In addition, the Halocynthia roretzi mitochondrial genome encodes an additional tRNA gene with the anticodon U*CU that is thought to enable the use of AGA or AGG codons for glycine and the gene has been shown to be transcribed in vivo (Kondow et al. 1999, Yokobori et al. 1999). Alternative initiation codons: ATA, GTG and TTG (Yokobori et al. 1999). ATT is the start codon for the CytB gene in Halocynthia roretzi (Gissi and Pesole, 2003) 14. The Alternative Flatworm Mitochondrial Code (transl_table=14)
AAs = FFLLSSSSYYY*CCWWLLLLPPPPHHQQRRRRIIIMTTTTNNNKSSSSVVVVAAAADDEEGGGG Starts = -----------------------------------M---------------------------- Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 14 Standard AAA Asn N Lys K AGA Ser S Arg R AGG Ser S Arg R UAA Tyr Y Ter * UGA Trp W Ter * Systematic Range:Platyhelminthes (flatworms) and Nematoda (roundworms)
Comments:
15. Blepharisma Nuclear Code (transl_table=15)
AAs = FFLLSSSSYY*QCC*WLLLLPPPPHHQQRRRRIIIMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = -----------------------------------M---------------------------- Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 10 GStandard UAG Gln Q Ter * Systematic Range:Ciliata: Blepharisma (Liang and Heckman, 1993)
16. Chlorophycean Mitochondrial Code (transl_table=16)
AAs = FFLLSSSSYY*LCC*WLLLLPPPPHHQQRRRRIIIMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = -----------------------------------M---------------------------- Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 16 Standard TAG Leu L STOP Systematic Range:Chlorophyceae (Hayashi-Ishimaru et al. 1996. UAG is a sense codon in several chlorophycean mitochondria) and the chytridiomycete fungus Spizellomyces punctatus (Laforest et al. 1997. Mitochondrial tRNAs in the lower fungus Spizellomyces punctatus: tRNA editing and UAG 'stop' codons recognized as leucine).
21. Trematode Mitochondrial Code (transl_table=21)
AAs = FFLLSSSSYY**CCWWLLLLPPPPHHQQRRRRIIMMTTTTNNNKSSSSVVVVAAAADDEEGGGG Starts = -----------------------------------M---------------M------------ Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 21 Standard TGA Trp W STOP ATA Met M Ile AGA Ser S Arg AGG Ser S Arg AAA Asn N Lys Systematic Range:Trematoda: Ohama, T, S. Osawa, K. Watanabe, T.H. Jukes, 1990. J. Molec Evol. 30Garey, J.R. and D.R. Wolstenholme, 1989. J. Molec. Evol. 28: 374-387 329-332.
22. Scenedesmus obliquus mitochondrial Code (transl_table=22)
AAs = FFLLSS*SYY*LCC*WLLLLPPPPHHQQRRRRIIIMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = -----------------------------------M---------------------------- Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format
Differences from the Standard Code:Code 22 Standard TCA STOP * Ser TAG Leu L STOP Systematic Range:Scenedesmus obliquus: Nedelcu A, Lee RW, Lemieux C, Gray MW and Burger G. "The complete mitochondrial DNA sequence of Scenedesmus obliquus reflects an intermediate stage in the evolution of the green algal mitochondrial genome." Genome Res. 2000 Jun;10(6):819-31.
23. Thraustochytrium Mitochondrial Code (transl_table=23)
AAs = FF*LSSSSYY**CC*WLLLLPPPPHHQQRRRRIIIMTTTTNNKKSSRRVVVVAAAADDEEGGGG Starts = --------------------------------M--M---------------M------------ Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format This code has been created for the mitochondrial genome of the labyrinthulid Thraustochytrium aureum sequenced by the The Organelle Genome Megasequencing Program (OGMP). It is the similar to the bacterial code (trans_table 11) but it contains an additional stop codon (TTA) and also has a different set of start codons.
24. Pterobranchia mitochondrial code (transl_table=24)
AAs = FFLLSSSSYY**CCWWLLLLPPPPHHQQRRRRIIIMTTTTNNKKSSSKVVVVAAAADDEEGGGG Starts = ---M---------------M---------------M---------------M------------ Base1 = TTTTTTTTTTTTTTTTCCCCCCCCCCCCCCCCAAAAAAAAAAAAAAAAGGGGGGGGGGGGGGGG Base2 = TTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGGTTTTCCCCAAAAGGGG Base3 = TCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAGTCAG Click here to change format Differences from the Standard Code:Code 24 Standard AGA Ser S Arg R AGG Lys K Arg R UGA Trp W STOP * Code 24 has been created for the mitochondrial genome of Rhabdopleura compacta (Pterobranchia). The Pterobranchia are one of the two groups in the Hemichordata which together with the Echinodermata and Chordata form the three major lineages of deuterostomes. AUA translates to isoleucine in Rhabdopleura as it does in the Echinodermata and Enteropneusta while AUA encodes methionine in the Chordata. The assignment of AGG to Lys is not found elsewhere in deuterostome mitochondria but it occurs in some taxa of Arthropoda (Perseke et al. 2011). Code 24 shares with many other mitochondrial codes the reassignment of the UGA STOP to Trp, and AGG and AGA to an amino acid other than Arg. The initiation codons in Rhabdopleura compacta are ATG and GTG (Perseke et al. 2011).
Comments and questions to info@ncbi.nlm.nih.gov Credits: Andrzej (Anjay) Elzanowski, Jim Ostell, Detlef Leipe, Vladimir Soussov. |
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