Skip Navigation Links

Contact Information:

photo of David Mattson
Name:
Mattson, David
Title:
Research Wildlife Biologist
Discipline(s):
Biology
Research Station:
CPRS
Work Address:
USGS Southwest Biological Science Center, Colorado Plateau Research Station, P.O. Box 5614, Northern Arizona University, Flagstaff, AZ 86011-5614
Telephone Number:
(928)523-9080
Email Address:
david_mattson@usgs.gov or David.Mattson@nau.edu

Biographical Sketch:

David Mattson is a Research Wildlife Biologist with the U.S. Geological Survey stationed at its' Colorado Plateau Research Station in Flagstaff, Arizona. David received degrees in Forest Resource Management and Forest Ecology and a doctorate in Wildlife Resource Management from the University of Idaho. Dr. Mattson has studied large carnivores for the last 24 years, focusing on puma ecology and human-puma interactions in Arizona, and the conservation and behavioral ecology of grizzly bears in the Yellowstone ecosystem of Wyoming, Montana, and Idaho. He spent 14 years intensively studying grizzly bear foraging behavior and diet as well as ecological relations of foods the bears ate. These studies revealed details about a broad spectrum of bear behaviors, including their bedding, use of rub trees, consumption of dirt and earthworms, exploitation of red squirrels, pocket gophers, and meadow mice, and predation on elk, moose, and trout. More recently, Dr. Mattson has focused on conservation issues and broad-scale evaluations of habitat conditions. These studies have broached not only the details of human-large carnivore interactions, but also the social, political, and organizational dynamics that shape the policies and practices of carnivore conservation programs. His work has been featured in the journal Science and has been widely presented, including papers in Ecology, Conservation Biology, Biological Conservation, The Journal of Wildlife Management, and the Journal of Mammalogy, and invited talks at the Smithsonian, American Museum of Natural History, the American Institute of Biological Sciences, and International Conferences on Bear Research and Management.

Project List

  • A. Pumas of the Flagstaff Uplands - Conflict between pumas and humans is receiving increased attention in the interior western United States as both human and puma populations increase. This project focuses on puma-human relations and the behavioral ecology of pumas in the Flagstaff uplands of northern Arizona. Project objectives include (i) description of puma movements and activity patterns, (ii) description of puma prey composition and predation rates, (iii) development of a study area GIS that includes human-related features, (iv) development of individual-based habitat selection models, and (iv) development of models and maps that explain and predict probability of contact between humans and pumas. 2003, Daily Sun, Flagstaff, AZ (Seth Muller); collaboration among USGS, NPS, and Grand Canyon Trust on a puma research project in the Flagstaff area of Arizona.
  • B. Ecological Effects of Upland Free Water - Free-standing water is a major limiting factor for many species in the arid West. This project focuses on the availability of free-standing water and its effects on biological communities in arid ecosystems of Arizona and Utah. Project objectives include (i) mapping free-water availability, (ii) explaining free-water availability in terms of geologic, geomorphic, climatic, and hydrologic features, (iii) describing temporal-spatial patterns of free-water use by animal species, (iv) determining the importance of free-standing water to different animal species, and (iv) determining the direct and indirect effects of artificial water sources on arid ecosystem animal communities.
  • C. Inventory & Monitoring of Biological Resources - The National Park Service has been charged with inventorying and monitoring its' biological resources. This project focuses on providing assistance to the National Park Service in designing and evaluating its' inventory and monitoring program for the Southern and Northern Colorado Plateau Networks. Assistance has taken the form of developing and writing planning, developing and evaluating protocols, and contributing to the selection of vital signs.
  • D. Conservation of Large Carnivores - Management of large carnivores embodies challenges that are common to the management of natural resources worldwide. This project focuses on facets of human psycho- sociology and behavior that affect conservation and management of large carnivores in North America. Project researchers map social processes and participant identities to analyze and explain decision-making related to different phases of formal and informal policy processes, considering factors such as political geography and organizational behavior. 2000-2001, WildFutures/Earth Island Institute (John de Graaf); conservation of grizzly bears for film "On Nature's Terms." 2000-2001, Time magazine and CNN (Bill Campbell); grizzly bear conservation. 1999, Audubon magazine (Todd Wilkinson); management and conservation of grizzly bears; featured in article in July-August 1999 issue. 1998, Washington Post (Tom Kenworthy); grizzly bear science and conservation in the Yellowstone ecosystem. 1998, New York Times (James Robbins); grizzly bear science and conservation in the Yellowstone ecosystem.
  • E. Modeling Habitat Suitability for Grizzly Bears - Models that predict source and restoration areas for extant and extirpated populations are potentially valuable to designing management that assures the long-term persistence of grizzly bears. This project focuses on developing models that explain and predict the extent and location of restoration and population source areas for grizzly bears, using varied data, varied study areas, information-based methods, and standards of sufficiency. These models are applied to existing and potential grizzly bear range throughout western North America. 2004, USA Today, (Tom Kenworthy); results of a paper published in Ursus that investigated habitat conditions for grizzly bears in the Cabinet-Yaak region of Montana-Idaho. 2003, Science Times of the New York Times (Jim Robbins); results of a paper published in Ursus that investigated habitat conditions for grizzly bears in the Cabinet-Yaak region of Montana-Idaho. 2003, Focus West, Public Broadcasting System (Jeff O'Gara); factors affecting the distribution of grizzly bears in the western United States. 2002, Billings Gazette, Billings, MT (Mike Stark); results of a paper published in Conservation Biology that investigated extirpations of grizzly bears in the western United States. 2002, Environmental News Network (Cat Lazaroff); results of a paper published in Conservation Biology investigating extirpations of grizzly bears in the western United States. 2002, National Parks & Conservation Association magazine (Todd Wilkinson); results of a paper published in Conservation Biology investigating extirpations of grizzly bears and projections of conditions for grizzly bear with global climate change. 2001, Audubon magazine (Todd Wilkinson); results of an article in Conservation Biology that investigated extirpations of grizzly bears in the western United States. 2000, Science magazine (Bee Wuethrich); umbrella effects; "When protecting one species hurts another." Science 289: 383, 385. 2000, Science Times of the New York Times (Mark Durer); reintroduction of grizzly bears into the Selway-Bitterroot ecosystem. 2000, The Denver Post (Kit Minnkler); reintroduction of grizzly bears into the Selway-Bitterroot ecosystem. 1999, The Spokesman-Review, Spokane, WA (Ken Olsen); results of a population viability analysis for grizzly bear in the contiguous United States and prospects for restoration of grizzly bears in central Idaho. 1999, High Country News (Juniper Davis); the historical importance of salmon to grizzly bears in Idaho. 1999, Idaho Statesman, Boise, ID (Rocky Barker); grizzly bear restoration in central Idaho. 1999, Toronto Globe & Mail (Anne MacAlroy); grizzly bear conservation along the U.S./Canadian border. 1999, Los Angeles Times (Frank Clifford); conservation issues along the Continental Divide.
  • F. Diet and Behavior of Grizzly Bears - Grizzly bears in the Yellowstone ecosystem of Wyoming, Montana, and Idaho were subjects of a unique long-duration and integrated study that included detailed investigations of diet and behavior during 1977-1996. This project focuses on analyzing data from this study, framed in terms of ecological theory, and related to local, regional, and global management of grizzly and brown bears. Different behaviors associated with bedding, tree rubbing, or the exploitation of specific foods are described and explained. Use of key foods is also related to movement and reproductive success of individual bears. 2003, Living Machine, National Geographic TV (Tierney Thys); grizzly bear consumption of carrion and wasps. 2003, German Public Radio (Ingrid Krölle); grizzly bear foods and foraging behavior and the ecological and cultural importance of grizzly bears. 2003, Nature, Public Broadcasting Corporation (Shane Moore); human-grizzly bear relations and the ecology of grizzly bears in the Yellowstone ecosystem and their future prospects. 2003, Billings Gazette, Billings, MT (Mike Stark); bark beetle-caused mortality among whitebark pine and consequences to grizzly bears in the Yellowstone ecosystem. 2003, The Denver Post (Theo Stein), Time magazine (Pat Clawson) and Defenders magazine (Dan Chasen); ecology of grizzly bears in the Yellowstone ecosystem and their future prospects. 2003, Casper Star Tribune, Casper, WY (Brodie Farguhar); ecology of grizzly bears in the Yellowstone ecosystem and their future prospects. 2002, Ventura County Star (John Krist), Outdoor Life (Andrew McKean), Billings Gazette (Mike Stark), and Los Angeles Times (Deborah Schoch); ecology and management of Yellowstone grizzly bears. 2001, Bozeman Chronicle, Bozeman, MT (Joan Haines); autecology of whitebark pine and threats and potential restoration strategies. 2000, Backpacker magazine (Erik Ness); implications of variability in grizzly bear food abundance to backpacker safety. 2000, British Broadcasting Corporation, Natural History Unit (Nigel Bean); Pleistocene history of North American bears. 2000, Los Angeles Times (Frank Clifford); status and future prospects of Yellowstone grizzly bears and their habitat. 2000, National Wildlife magazine (Gary Turbak); grizzly bear ecology and the status of grizzly bear foods. 1999, Environmental Review newsletter (Douglas Taylor); ecology and management of Yellowstone grizzly bears; featured interview in the August 1999 (Volume 6[8]) issue. 1999 & 2000, British Broadcasting Corporation, 'The Natural World' (Jonathan Keeling); brown bears of the world and grizzly bears in Yellowstone. 1998, National Geographic (Jennifer Steinberg); relations between grizzly bears and cutthroat trout in the Yellowstone ecosystem.

Publications, Maps, and other Products

  1. Mattson, D. J., K. L. Byrd, M. B. Rutherford, S. R. Brown, & T. W. Clark. 2006. Finding common ground in large carnivore conservation; Mapping contending perspectives. Environmental Science and Policy 9:392-405. [Journal Article]
  2. Wilson, S. M., M. J. Madel, D. J. Mattson, J. M. Graham, T. Merrill. 2006. Landscape conditions predisposing grizzly bears to conflicts on private agricultural lands in the western USA. Biological Conservation 130:47-59. [Journal Article]
  3. Mattson, David. 2005. Upland Free Water in the Southwest. A Research Program for Managment of a Critical Resource in the Arid & Semi-Arid West. [Fact Sheet]
  4. Clark, T. W., & D. J. Mattson. 2005. Making carnivore management programs more effective: a guide for decision making. Coexisiting with large Carnivores, pp 271-276, T. W. Clark, M. B. Rutherford, & D. Casey, editors. Island Press, Washington, D.C. [Book Chapter]
  5. Mattson, D. J., S. Herrero & T. Merrill. 2005. Are black bears a factor in the restoration of North American grizzly bear populations. Ursus 16:11-30. [Journal Article]
  6. Wilson, S. M., M. J. Madel, D. J. Mattson, J. M. Graham, J. A. Burchfield, & J. M. Belsky. 2005. Natural landscape features, human-related attractants, and conflict hotspots: a spatial analysis of human-grizzly bear conflicts. Ursus 16:117-129. [Journal Article]
  7. Mattson, D. J., S. R. Podruzny, & M. A. Haroldson. 2005. Consumption of pondweed rhizomes by Yellowstone grizzly bears. Ursus 16:41-46. [Journal Article]
  8. Mattson, D. 2005. Seeing the elephant: holistic intelligence for solving wildlife-related problems. Proceedings of the Conference on Interdisciplinary Research and Management in Mountain Areas, pp 53-56, L. Taylor & P. Murphy, editors. The Banff Centre, Banff, AB, Canada. [Proceedings Chapter]
  9. Mattson, D., J. Hart, & T. Arundel. 2005. Cougar predation in the Flagstaff uplands: preliminary results from July 2003-May 2005. Proceedings of the 8th Mountain Lion Workshop, pp 158-169. Washington Department of Fish & Game, Olympia WA. [Proceedings Chapter]
  10. Mattson, D. J. 2004. Consumption of voles and vole food caches by Yellowstone grizzly bears; exploratory analyses. Ursus 15:218-226. [Journal Article]
  11. Mattson D.J., T. Merrill. 2004. A model-based appraisal of habitat conditions for grizzly bears in the Cabinet-Yaak region of Montana and Idaho. Ursus 15 : 78-91. [Journal Article]
  12. Mattson, D.J. 2004. Consumption of pocket gophers and their food caches by grizzly bears. Journal of Mammalogy 85 : 000-000. [Journal Article]
  13. Johnson M., J. Holmes, D. Mattson, L. Thomas, N. Tancreto. 2004. Summary of faunal populations and communities workshop April 6-7, 2004, Northern Arizona University , Flagstaff , Arizona NPS, Southern Colorado Plateau I&M Network. U.S. National Park Service, Southern Colorado Plateau I&M Network, Flagstaff , Arizona . 10pp. [Technical Report]
  14. Mattson, D.J. 2004. Living with fierce creatures? An overview and models of mammalian carnivore conservation. Pages 00-00 in N. Fascione, A. Delach & M. Smith, editors. Predators and people: from conflict to conservation . Island Press. 28pp. [Book Chapter]
  15. Mattson D.J., K. Barber, R. Maw, R. Renkin. 2004. Coefficients of productivity for Yellowstone's grizzly bear habitat. U.S. Geological Survey, Biological Science Report USGS/BRD/BSR-2002-0007. 76pp. [Monograph]
  16. Mattson D.J., J.V. Hart, P. Beier, J. Millen-Johnson. 2003. A conceptual model and appraisal of existing research related to interactions between humans and pumas. Page 104 in S.A. Becker, D.D. Bljornlie, F.G. Lindzey, & D. S. Moody, editors. Porceedings of the Seventh Mountain Lion Workshop. Wyoming Game & Fish Department, Lander, WY. [Abstract]
  17. Merrill T., D.J. Mattson. 2003. The extent and location of habitat biophysically suitable for grizzly bears in the Yellowstone region. Ursus 14 : 171-187. [Journal Article]
  18. Mattson D.J. 2003. Conservation of mountain carnivores: living with fierce creatures? . In K. Martin & D.S. Hik, editors. Mountain Science Highlights. [Web Page]
  19. Mattson D.J. 2003. Foot loadings and pad and track widths of Yellowstone grizzly bears. Western North American Naturalist 63 : 72-79. [Journal Article]
  20. Mattson D.J. 2003. Thoughts on designing a monitoring program for the Southern Colorado Plateau Network (SCPN) National Park units. For US National Park Service Southern Colorado Plateau Network, Flagstaff , AZ. USGS Southwest Biological Science Center . 4pp. [Technical Report]
  21. Mattson D.J. 2003. Some thoughts on evaluating the Yellowstone Grizzly Bear Cumulative Effects Model. For Interagency Grizzly Bear Study Team, Bozeman , MT. USGS Southwest Biological Science Center. 3pp. [Technical Report]
  22. Mattson D.J. 2003. "Conservation is like warfare:" phantom common ground in the grizzly bear case. For Yale School of Forestry & Environmental Studies, Seminar on Society & Natural Resources (F&ES 746). 7pp. [Abstract]
  23. Mattson D.J., S.R. Podruzny, M.A. Haroldson. 2002. Consumption of fungal sporocarps by Yellowstone grizzly bears. Ursus 13 : 159-168. (Manuscript chosen as an exemplar for authors submitting manuscripts to Ursus). [Journal Article]
  24. Mattson D.J., J. Hart, T. Arundel. 2002. Cougars of the Flagstaff uplands. USGS Southwest Biological Science Center , Flagstaff , AZ. 2pp. [Fact Sheet]
  25. Mattson D.J., M.G. French, S.P. French. 2002. Consumption of earthworms by Yellowstone grizzly bears. Ursus 13 : 153-158.( Abstract chosen as an exemplar for technical writing class, New Mexico State University ). [Abstract]
  26. Mattson D.J., T. Merrill. 2002. Extirpations of grizzly bears in the contiguous United States, 1850-2000. Conservation Biology 16 : 1123-1136. ( Paper featured in press release by Conservation Biology and solicited for inclusion in eForum on Biocomplexity & Conservation ). [Journal Article]
  27. Mattson D.J. 2002. Consumption of wasps and bees by Yellowstone grizzly bears. Northwest Science 76 : 166-172. [Journal Article]
  28. Mattson D.J. 2002. An approach to selecting vital signs for the Colorado Plateau National Park Service inventory and monitoring program. For US National Park Service Northern Colorado Plateau Network , Moab , UT. USGS Forest & Rangeland Ecosystem Science Center , Colorado Plateau Field Station. 7pp. [Technical Report]
  29. Green G. I., D. J. Mattson. 2002. Tree rubbing by Yellowstone grizzly bears Ursus arctos. Wildlife Biology 9:1-9. [Journal Article]
  30. Hendee J.C., D.J. Mattson. 2002. Wildlife in wilderness: a North American and international perspective. Pages 321-349 in J.C. Hendee & C.P. Dawson, editors. Wilderness management: stewardship and protection of resources and values. 3 rd edition . Fulcrum Publishing, Golden, CO. [Book Chapter]
  31. Reinhart D.P., M. Haroldson, D.J. Mattson, K.A. Gunther. 2001. Effects of exotic species on Yellowstone 's grizzly bears. Western North American Naturalist 61 : 277-288. [Journal Article]
  32. Mattson D.J., K.C. Kendell, D.P. Reinhart. 2001. Whitebark pine, grizzly bears and red squirrels. Pages 121-136 in D.F. Tomback, S.F. Arno, & R.E. Keane, editors. Whitebark pine communities: ecology and restoration . Island Press, Washington , D.C. [Book Chapter]
  33. Mattson D.J. 2001. The effects of fragmentation, edges and habitat loss on wildlife: a perspective for mountain wildlife. Pages 191-200 in L. Taylor & A. Ryall, editors. Human use management in mountain areas. The Banff Centre, Banff , AB. [Book Chapter]
  34. Mattson D.J. 2001. Myrmecophagy by Yellowstone grizzly bears. Canadian Journal of Zoology 79 : 779-793. [Journal Article]
  35. Mattson D.J. 2001. Comments on ecological effects of the four-wheel-drive route in Salt Creek, Canyonlands National Park, Utah. For Southeast Utah Group National Parks & Monuments, Moab , UT. USGS Forest & Rangeland Ecosystem Science Center , Colorado Plateau Field Station. 14pp. [Technical Report]
  36. Clark T.W., D.J. Mattson, R.P. Reading, Miller. 2001. Interdisciplinary problem solving in carnivore conservation: an introduction. Pages 223-240 in J.L. Gittleman, S.M. Funk, D. Macdonald, & R.K. Wayne, editors. Carnivore conservation . Cambridge University Press, Cambridge , U.K. [Book Chapter]
  37. Mattson D.J. 2000. Grizzly bears. Pages 66-71 in R.P. Reading & B. Miller, editors. Endangered animals: a reference guide to conflicting issues. Greenwood Press, Westport, CT. [Book Chapter]
  38. Mattson, D.J. 2000. Causes and consequences of dietary differences among Yellowstone grizzly bears ( Ursus arctos ). Ph.D. Dissertation, University of Idaho , Moscow , ID. 173 pp. [Thesis/Dissertation]
  39. Mattson, D.J. 2000. Managing whitebark pine for grizzly bears: preliminary recommendations. For Interagency Grizzly Bear Study Team, Bozeman , MT. USGS Forest & Rangeland Ecosystem Science Center , Colorado Plateau Field Station. 3pp. [Technical Report]
  40. Drost C., D.J. Mattson, M.J. Johnson, A. Cully, M. Bogan, E. Nowak, T. Persons, J. Spence, K. Thomas, M. Stuart. 2000. Biological inventory of National Park areas on the southern Colorado Plateau. For US National Park Service Southern Colorado Plateau Network. Colorado Plateau Cooperative Ecosystem Studies Unit and USGS Colorado Plateau Field Station, Flagstaff , AZ. 209pp. (INVITED technical plan; rated second-best inventory plan nationwide ). [Technical Report]
  41. Podruzny S.R., D.P. Reinhart, D.J. Mattson. 1999. Fire, red squirrels, whitebark pine, and Yellowstone grizzly bears. Ursus 11 : 131-138. [Journal Article]
  42. Mattson D.J., G.I. Green, R.A. Swalley. 1999. Geophagy by Yellowstone grizzly bears. Ursus 11 : 109-116. [Journal Article]
  43. Scott J.M., E.A. Norse, H. Arita, A. Dobson, J.A. Estes, M. Foster, B. Gilbert, D.B. Jenson, R.L. Knight, D. Mattson, M.E. Soulé. 1999. The issues of scale in selecting and designing biological reserves. Pages 19-37 in M.E. Soulé & J. Terborgh, editors. Continental conservation . Island Press, Washington , D.C. [Book Chapter]
  44. Pease C.M., D.J. Mattson. 1999. Demography of the Yellowstone grizzly bears. Ecology 80 : 957-975. (Paper featured in press release by Ecology and in an article by Science). [Journal Article]
  45. Merrill T., D.J. Mattson, R.G. Wright, H.B. Quigley. 1999. Defining landscapes suitable for restoration of grizzly bears Ursus arctos in Idaho. Biological Conservation 87 : 231-248. [Journal Article]
  46. Dobson A., K. Ralls, M. Foster, M.E. Soulé, D. Simberloff, D. Doak, J.A. Estes, L.S. Mills, D. Mattson, R. Dirzo, H. Arita, S. Ryan, E.A. Norse, R.F. Noss, D. Johns. 1999. Corridors: reconnecting fragmented landscapes. Pages 129-170 in M.E. Soulé & J. Terborgh, editors. Continental conservation . Island Press, Washington , D.C. [Book Chapter]
  47. Merrill T., D.J. Mattson. 1998. Defining grizzly bear habitat in the Yellowstone to Yukon. Pages 103-111 in A. Harvey, editor. A sense of place: issues, attitudes and resources in the Yellowstone to Yukon ecoregion. Yellowstone to Yukon Conservation Initiative, Canmore , AB. [Journal Article]
  48. Merrill T., D.J. Mattson. 1998. Land cover structure of Yellowstone to Yukon. Pages 27-29 in A. Harvey, editor. A sense of place: issues, attitudes and resources in the Yellowstone to Yukon ecoregion. Yellowstone to Yukon Conservation Initiative, Canmore, AB. [Publication]
  49. Mattson D.J. 1998. Changes in the mortality of Yellowstone 's grizzly bears. Ursus 10 : 129 ? 138. [Journal Article]
  50. Mattson D.J. 1998. Diet and morphology of extant and recently extinct northern bears. Ursus 10 : 479-496. [Journal Article]
  51. Mattson, D.J. 1998. Coefficients of productivity for Yellowstone's grizzly bear habitat. USGS Forest & Rangeland Ecosystem Science Center, Corvallis, OR. 85pp. [Technical Report]
  52. Mattson D.J. 1998. Research problem analysis: Yellowstone's grizzly bear research program. For Interagency Grizzly Bear Study Team, Bozeman , MT. USGS Biological Resources Division, Forest & Rangeland Ecosystem Science Center . 10pp. [Technical Report]
  53. Mattson D.J., D.P. Reinhart. 1997. Excavation of red squirrel middens by Yellowstone grizzly bears. Journal of Applied Ecology 34 : 926-940. [Journal Article]
  54. Mattson D.J. 1997. Wilderness-dependent wildlife: the large and the carnivorous. International Journal of Wilderness 3 : 34-38. [Journal Article]
  55. Mattson D.J. 1997. Sustainable grizzly bear mortality calculated from counts of females with cubs-of-the-year: an evaluation. Biological Conservation 81 : 103-111. [Journal Article]
  56. Mattson, D.J. 1997. Use of lodgepole pine cover types by Yellowstone grizzly bears. Journal of Wildlife Management 61 : 480-496. [Journal Article]
  57. Mattson D.J. 1997. Use of ungulates by Yellowstone grizzly bears Ursus arctos. Biological Conservation 81 : 161-177. [Journal Article]
  58. Green G.I., D.J. Mattson, J.M. Peek. 1997. Spring feeding on ungulate carcasses by grizzly bears in Yellowstone National Park. Journal of Wildlife Management 61 : 1040 ! 1055. [Journal Article]
  59. Mattson D.J. 1997. Selection of microsites by grizzly bears to excavate biscuitroots. Journal of Mammalogy 78 : 228-238. [Journal Article]
  60. Wright R. G., D.J. Mattson. 1996. The origin and purpose of national parks and protected areas. Pages 3-14 in R.G. Wright, editor. National parks and protected areas: their role in environmental protection. Blackwell Science, Cambridge, MA . [Book Chapter]
  61. Mills L.S., S.G. Hayes, C. Baldwin, M.J. Wisdom, J. Citta, D.J. Mattson, K. Murphy. 1996. Factors leading to different viability predictions for a grizzly bear data set. Conservation Biology 10 : 863-873. [Journal Article]
  62. Mattson D. J., S. Herrero, R.G. Wright, C.M. Pease. 1996. Designing and managing protected areas for grizzly bears: how much is enough. Pages 133-164 in R.G. Wright, editor. National Parks and protected areas: their role in environmental protection. Blackwell Science, Cambridge, MA. [Book Chapter]
  63. Mattson D.J., S. Herrero, R.G. Wright, C.M. Pease. 1996. Science and management of Rocky Mountain grizzly bears. Conservation Biology 10 : 1013-1025. [Journal Article]
  64. Mattson D.J., D.P. Reinhart. 1996. Indicators of red squirrel ( Tamiasciurus hudsonicus ) abundance in the whitebark pine zone. Great Basin Naturalist 56 : 272-275. [Journal Article]
  65. Mattson D.J. 1996. Ethics and science in natural resource agencies. BioScience 46 : 767-771. [Journal Article]
  66. Mattson D.J. 1996. Modeling grizzly bear habitat suitability in Idaho. GAP Analysis Program Bulletin 5 : 18-19. ( in Best of Gap ). [Publication]
  67. Mattson D.J., R.G. Wright, K. C. Kendall, C.J. Martinka. 1995. Grizzly bears. Pages 103-105 in E.T. LaRoe, G.S. Farris, C.E. Puckett, P.D. Doran & M.J. Mac, editors. Our living resources: a report to the nation on the distribution, abundance, and health of U.S. plants, animals, and ecosystems . U.S. National Biological Service, Washington , D.C. [Book Chapter]
  68. Mattson D. J., D.P. Reinhart. 1995. Influences of cutthroat trout ( Oncorhynchus clarki ) on behaviour and reproduction of Yellowstone grizzly bears ( Ursus arctos ), 1975-1989. Canadian Journal of Zoology 73 : 2072-2079. [Journal Article]
  69. Mattson D.J. 1995. The New World Mine and grizzly bears: a window on ecosystem management. Journal of Energy, Natural Resources & Environmental Law 15 : 267-293. [Journal Article]
  70. Mattson D.J. 1995. Book review - "Greater Yellowstone's future: prospects for ecosystem science, management, and policy.". Endangered Species Update 12 : 10. [Book Review]
  71. Mattson D.J., D.P. Reinhart, B.M. Blanchard. 1994. Variation in production and bear use of whitebark pine seeds in the Yellowstone area. Pages 205-220 in D.G. Despain, editor. Plants and their environments: proceedings of the first biennial scientific conference on the Greater Yellowstone Ecosystem. U.S. National Park Service Technical Report NPS/NRYELL/NRTR. [Technical Report]
  72. Mattson D.J., D.P. Reinhart. 1994. Bear use of whitebark pine seeds in North America. Pages 212-220 in W.C. Schmidt & F.-K. Holtmeier, compilers. Proceedings - international workshop on subalpine stone pines and their environment: the status of our knowledge . U.S. Forest Service General Technical Report INT-GTR-309 . [Technical Report]
  73. Mattson D.J., J.J. Craighead. 1994. The Yellowstone grizzly bear recovery program: uncertain information, uncertainty policy. Pages 101-129 in T.W. Clark, R.P. Reading & A. Clarke, editors. Endangered species recovery: finding the lessons, improving the process. Island Press, Washington , D.C. [Book Chapter]
  74. Mattson D.J., I. Revenko. 1993. Kamchatkan brown bears and Pinus pumila. Pages 48-49 in W. McCaughey & K.J. McDonald, compilers. Management of whitebark pine ecosystems - an international and regional perspective . U.S. Forest Service Intermountain Research Station, Ogden, UT. [Abstract]
  75. Mattson D.J. 1993. Background and proposed standards for managing grizzly bear habitat security in the Yellowstone ecosystem. U.S. National Biological Survey, University of Idaho Cooperative Park Studies Unit, Moscow. 17pp. [Technical Report]
  76. Reinhart D.P., D.J. Mattson. 1992. Grizzly bear and black bear habitat use in the Cooke City, Montana, area, 1990-1991. U.S. National Park Service, Interagency Grizzly Bear Study Team, Bozeman, MT. 31pp. [Technical Report]
  77. Mattson D.J., R.R. Knight, B.M. Blanchard. 1992. Cannibalism and predation on black bears by grizzly bears in the Yellowstone ecosystem, 1975-1990. Journal of Mammalogy 73 : 22-425. [Journal Article]
  78. Mattson D.J., B.M. Blanchard, R.R. Knight. 1992. Yellowstone grizzly bear mortality, human habituation, and whitebark pine seed crops. Journal of Wildlife Management 56 : 432-442. [Journal Article]
  79. Mattson D.J., R.R. Knight. 1992. Spring bear use of ungulates in the Firehole River drainage of Yellowstone National Park. Pages 5.93-5.120 in J.D. Varley & W.G. Brewster, editors. Wolves for Yellowstone? A report to the United States Congress, volume IV, research and analysis. U.S. National Park Service, Yellowstone National Park, WY. [Publication]
  80. Mattson D.J. 1992. The Yellowstone experience: "between a rock and a hard place.". Pages 23-30 in R.K. McCann, editor. Grizzly bear management workshop proceedings. Environment Canada, Parks Service, Revelstoke, BC. [Publication]
  81. Blanchard B.M., R.R. Knight, D.J. Mattson. 1992. Distribution of Yellowstone grizzly bears during the 1980s. American Midland Naturalist 128 : 332-338. [Journal Article]
  82. Mattson D.J., C.M. Gillin, S.A. Benson, R.R. Knight. 1991. Bear feeding activity at alpine insect aggregation sites in the Yellowstone ecosystem. Canadian Journal of Zoology 69 : 430-2435. [Journal Article]
  83. Mattson D.J., B.M. Blanchard, R.R. Knight. 1991. Food habits of Yellowstone grizzly bears, 1977-1987. Canadian Journal of Zoology 69 : 1619-1629. [Journal Article]
  84. Mattson D.J., M. W. Reid. 1991. Conservation of the Yellowstone grizzly bear. Conservation Biology 5 : 364-372. [Journal Article]
  85. Mattson D.J., R.R. Knight. 1991. Application of cumulative effects analysis to the Yellowstone grizzly bear population. U.S. National Park Service, Interagency Grizzly Bear Study Team Report 1991C, Bozeman, MT. 8pp. [Technical Report]
  86. Reinhart D.P., D.J. Mattson. 1990. Bear use of cutthroat trout spawning streams in Yellowstone National Park. International Conference on Bear Research & Management 8 : 343-350. [Journal Article]
  87. Reinhart D.P., D.J. Mattson. 1990. Red squirrels in the whitebark pine zone. Pages 256-263 in W.C. Schmidt & K.J. McDonald, compilers. Proceedings - symposium on whitebark pine ecosystems: ecology and management of a high-mountain resource. U.S. Forest Service General Technical Report INT-270. [Technical Report]
  88. Mattson D.J., D. P. Reinhart. 1990. Whitebark pine on the Mount Washburn massif, Yellowstone National Park. Pages 106-117 in W.C. Schmidt & K.J. McDonald, compilers. Proceedings - symposium on whitebark pine ecosystems: ecology and management of a high-mountain resource. U.S. Forest Service General Technical Report INT-270. [Technical Report]
  89. Mattson D.J., C. Jonkel. 1990. Stone pines and bears. Proceedings - symposium on whitebark pine ecosystems: ecology and management of a high-mountain resource. U.S. Forest Service General Technical Report INT-270. Pages 223-236 in W.C. Schmidt & K.J. McDonald, compilers. [Proceedings]
  90. Mattson D.J. 1990. Human impacts on bear habitat use. International Conference on Bear Research & Management 8 : 35-56. [Journal Article]
  91. Mattson D.J., R.R. Knight. 1989. Evaluation of grizzly bear habitat using habitat and cover type classifications. Pages 135-143 in D.E. Ferguson, P. Morgan & F.D. Johnson, compilers. Proceedings - land classifications based on vegetation: applications for resource management. U.S. Forest Service General Technical Report INT-257. [Proceedings]
  92. Mattson D., G. Green, J. Henry. 1988. Dynamics of ungulate carcasses and their use by bears on ungulate winter ranges. Page 18 in F. Singer, compiler. First annual meeting of research and monitoring on Yellowstone 's northern range. U.S. National Park Service, Yellowstone National Park, WY. [Abstract]
  93. Mattson D.J., R.R. Knight, B.M. Blanchard. 1987. The effects of developments and primary roads on grizzly bear habitat use in Yellowstone National Park, Wyoming. International Conference on Bear Research & Management 7 : 259-273. [Journal Article]
  94. Weaver J., R. Esacno, D. Mattson, T. Puchlerz. 1986. Cumulative effects process for the Yellowstone ecosystem. Pages 234-246 in G. Contreras & K. Evans, compilers. Proceedings - grizzly bear habitat symposium. U.S. Forest Service General Technical Report INT-207. [Proceedings]
  95. Picton H., D. Mattson, B. Blanchard, R. Knight. 1986. Climate, carrying capacity, and the Yellowstone grizzly bear. Pages 129-135 in G. Contreras & K. Evans, compilers. Proceedings - grizzly bear habitat symposium. U.S. Forest Service General Technical Report INT-207. [Proceedings]
  96. Mattson D.J., R. R. Knight, B.M. Blanchard. 1986. Derivation of habitat component values for the Yellowstone grizzly bear. Pages 222-229 in G. Contreras & K. Evans, compilers. Proceedings - grizzly bear habitat symposium. U.S. Forest Service General Technical Report INT-207. [Proceedings]
  97. Mattson D.J., D. G. Despain. 1985. Grizzly bear habitat component mapping handbook for the Yellowstone ecosystem. U.S. National Park Service, U.S. Forest Service, Mammoth, WY. 37pp. [Technical Report]
  98. Mattson D.J. 1984. Classification and environmental relationships of wetland vegetation in central Yellowstone National Park. M.S. Thesis, University of Idaho, Moscow. 409pp. (Thesis). [Thesis/Dissertation]
  99. Brunsfeld S.J., L. Ashland, C.R. Hatch, D. J. Mattson, A.L. Merkel, M. Sargent, G.H. Savage. 1982. Biomass resources in 66 African, Asian, and Latin American countries. University of Idaho Forestry, Wildlife & Range Experiment Station, Contribution 231. [Technical Report]