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Printed in the United States of America for The Wildlife Society by Allen Press, Inc., Lawrence, Kansas

All rights in this book are reserved. No part of the book may be used or reproduced in any manner whatsoever without written permission except in the case of nonprofit educational reproduction, use by agencies of the U.S. Government, or brief quotations embodied in critical articles and review. For information, address: The Wildlife Society, Inc., 5410 Grosvenor Lane Bethesda, Maryland 20814

Mention of trade names does not imply endorsement of the product.

This book is the fifth in a series on wildlife techniques published by The Wildlife Society

Editor, Henry S. Mosby
     Manual of Game Investigational Techniques
        (1) First Edition-May 1960
             Second Printing-February 1961
     Wildlife Investigational Techniques
        (2) Second Edition-May 1963
             Second Printing (Revised)-March 1965
             Third through Sixth Printing-March 1966 to September 1968

Editor, Robert H. Giles, Jr.
     Wildlife Management Techniques
        (3) Third Edition-June 1969
             Second Printing (Revised)-January 1971
             Third Printing-May 1972

Editor, Sanford D. Schemnitz
     Wildlife Management Techniques Manual
        (4) Fourth Edition-September 1980

Editor, Theodore A. Bookhout
     Research and Management Techniques for Wildlife and Habitats
        (5) Fifth Edition-January 1994

Suggested citation formats:

Bookhout, T. A., Editor. 1994. Research and management techniques for wildlife 
     and habitats. Fifth ed. The Wildlife Society, Bethesda, Md. 740pp.

Johnson, D. H. 1994. Population analysis. Pages 419-444 in T. A. Bookhout, ed. 
     Research and management techniques for wildlife and habitats. Fifth ed. 
     The Wildlife Society, Bethesda, Md.

This book was produced on the Penta DeskTopPro_/UX^® and output to an AGFA SelectSet 7000 imagesetter. The text is Adobe Times Roman. The text paper is 50 pound Simpson Offset (50/10 recycled). The Roxite cloth cover was printed by Allen Press, Inc. and the case binding was done by Prizma Industries, Denver, Colorado. This book was printed on a Hantscho full-sized waterless web press by Allen Press, Inc.

ISBN 0-933564-10-4
Library of Congress Catalog Card Number: 93-61624