Jon E Keeley

USGS Sequoia and Kings Canyon Field Station
47050 Generals Highway #4
Three Rivers, CA 93271-9651
Phone: (559) 565-3170
Fax: (559) 565-3177
Email: jon_keeley@usgs.gov

• Adaptation
• Biodiversity
• Biogeography
• Chaparral Ecology
• Coastal Sage Ecology
• Community ecology
• Conservation biology
• Ecotone
• Endangered habitat
• Environmental impacts
• Evolution
• Extinction
• Fire ecology
• Forest ecology
• Global change biology
• Grassland Ecology
• Historical Reconstruction
• Invasive species ecology
• Landscape ecology
• Landscape patterns
• Natural history
• Phylogeny
• Physiological plant ecology
• Plant ecology
• Plant taxonomy
• Population biology
• Restoration ecology
• Species identification
• Species interactions
• Species introduction
• Species inventory
• Species/Population management
• Systematics
• Vegetation classification
• Vernal Pool Conservation
• Vernal Pool Ecology

• Ph.D., Botany, University of Georgia, Athens, GA 1977
• M.S., Biology, San Diego State University, San Diego, CA 1973
• B.S., Biology, San Diego State University, San Diego, CA 1971

• American Association for the Advancement of Science
• American Association of Plant Taxonomists
• British Ecological Society
• California Botanical Society
• Ecological Society of America
• International Association for Vegetation Science
• International Association of Wildland Fire
• Society for Conservation Biology
• Southern California Botanists

• Station Leader, U.S. Geological Survey, Sequoia-Kings Canyon, Jul 1998-Present
• Program Director, National Science Foundation, Washington, D.C., Sep 1997-Jul 1998
• Professor of Biology, Occidental College, Los Angeles, CA, Jul 1988-Sep 1998
• Visiting Professor, University of Cape Town, South Africa, Aug 1990-Nov 1990
• Department Chair, Occidental College, Los Angeles, CA, Jul 1982-Jun 1988
• Associate Professor, Occidental College, Los Angeles, CA, Jul 1982-Jun 1988
• Assistant Professor, Occidental College, Los Angeles, CA, Sep 1977-Jun 1982

• Bowman, D.M.J.S., J.K. Balch, P. Artaxo, W.J. Bond, J.M. Carlson, M.A. Cochrane, C.M. D’Antonio, R.S. DeFries, J.C. Doyle, S.P. Harrison, F.H. Johnston, J.E. Keeley, M.A. Krawchuk, C.A. Kull, J.B. Marston, M.A. Moritz, I.C. Prentice, C.I. Roos, A.C. Scott, T.W. Swetnam, G.R. van der Werf, and S.J. Pyne. 2009. Fire in the Earth system. Science 324:481–484.[Journal Article]
• Keeley, J.E. 2009. Fire intensity, fire severity and burn severity: A brief review and suggested usage. International Journal of Wildland Fire 18:116-126.[Journal Article]
• Keeley, J.E. and P.H. Zedler. 2009. Large, high intensity fire events in southern California shrublands: debunking the fine-grained age-patch model. Ecological Applications 19:69-94.[Journal Article]
• Keeley, J.E. and P.H. Zedler. 2009. Large, high intensity fire events in southern California shrublands: debunking the fine-grained age-patch model. Ecological Applications 19:69-94. (APPENDICES)[Journal Article]
• Keeley, J.E., G.H. Aplet, N.L. Christensen, S.G. Conard, E.A. Johnson, P.N. Omi, D.L. Peterson, and T.W. Swetnam. 2009. Ecological foundations for fire management in North American forest and shrubland ecosystems. USDA Forest Service, Pacific Northwest Research Station, General Technical Report, PNW-GTR-779. 92 p.[Technical Report]
• Schwilk, D.W., J.E. Keeley, E.E. Knapp, J. McIver, J.D. Bailey, C.J. Fettig, C.E. Fiedler, R.J. Harrod, J.J. Moghaddas, K.W. Outcalt, C.N. Skinner, S.L. Stephens, T.A. Waldrop, D.A. Yassey, and A. Youngblood. 2009. The National Fire and Fire Surrogate Study: Effects of alternative fuel reduction methods on forest vegetation structure and fuels. Ecological Applications 19:285-304.[Journal Article]
• Stephens, S.L., J.J Moghaddas, C. Edminster, C.E. Fiedler, S. Hasse, M. Harrington, J.E. Keeley, E. Knapp, J.D. McIver, K. Metlen, C. Skinner, and A. Youngblood. 2009. Fire and fire surrogate treatment effects on vegetation structure, fuels, and potential fire behavior and severity from six western United States coniferous forests. Ecological Applications 19:305-320.[Journal Article]
• Engles, D.L., J.O. Sickman, C.M. Moore, A.M. Esperanza, J.M. Melack, and J.E. Keeley. 2008. Biogeochemical legacy of prescribed fire in a giant sequoia-mixed conifer forest: A 16-year record of watershed balances. Journal of Geophysical Research – Biogeosciences 113: G01014[Journal Article]
• Keane, R.E., J.K. Agee, P. Fule, J.E. Keeley, C. Key, S.G. Kitchen, R. Miller, and L.A. Schulte. 2008. Ecological effects of large fires on US landscapes: benefit or catastrophe? International Journal of Wildland Fire 17:696-712.[Journal Article]
• Keeley, J.E. 2008. Relative impact of weather vs. fuels on fire regimes of coastal California, pp. 263-269. Fire Conference 2002: Managing Fire and Fuels in the Remaining Wildlands and Open Spaces of the Southwestern United States. USDA Forest Service, Pacific Southwest Research Station, General Technical Report PSW-GTR-189.[Technical Report]
• Keeley, J.E. and P.J. van Mantgem. 2008. Community ecology of seedlings, pp. 253- 270. In M.A. Leak, V.T. Parker, and R.L. Simpson, editors. Seedling ecology and evolution. Cambridge University Press, Cambridge, U.K.[Book Chapter]
• Keeley, J.E., T. Brennan, and A.H. Pfaff. 2008. Fire severity and ecosystem responses following crown fires in California shrublands. Ecological Applications 18:1530-1546. [Journal Article]
• Parker, V.T., M.C. Vasey, and J.E. Keeley. 2008. Taxonomic revisions in the genus Arctostaphylos. Madroño 54:148-155.[Journal Article]
• Pincetl, S., P.W. Rundel, J.C. DeBlasio, D. Silver, T. Scott, J.E. Keeley and R. Halsey. 2008. It’s the land use not the fuels: Fires and land development in southern California. Real Estate Review 37(1):25-42.[Journal Article]
• Keeley, J. E., A Critical Assessment of the Burning Index Used in Fighting Wildland Fires (August 2007) [Publication Brief]
• Keeley, J. E., Fire Dynamics of Foothill Pine in the Southern Sierra Nevada (February 2007)[Publication Brief]
• Keeley, J. E., Human Influence on California Fire Regimes (July 2007)[Publication Brief]
• Keeley, J. E., Prescribed Fire Effects on Cheatgrass Persistence in a Sierra Nevada Ponderosa Pine Forest, (February 2007) [Publication Brief]
• Keeley, J. E., Role of Burning Season on Understory Vegetation in a Mixed Conifer Forest (March 2007) [Publication Brief]
• Keeley, J. E., The Role of Fuel Breaks in the Invasion of Nonnative Plants (April 2007) [Publication Brief]
• Keeley, J.E. 2007. Appropriate postfire management for the 2007 Griffith park Fire. Newsletter of the Los Angeles & San Gabriel Rivers Watershed Council 11(1):3-5.[Journal Article]
• Keeley, J.E. 2007. Chaparral and fire. Fremontia 35(4):16-21.[Journal Article]
• Keeley, J.E. and T.W. McGinnis. 2007. Impact of prescribed fire and other factors on cheatgrass persistence in a Sierra Nevada ponderosa pine forest. International Journal of Wildland Fire 16:96-106.[Journal Article]
• Keeley, J.E., and F.W. Davis. 2007. Chaparral. Pages 339-366 in: M.G. Barbour, T. Keeler-Wolf, and A.A. Schoenherr (eds.), Terrestrial Vegetation of California, 3rd edition. University of California Press, Los Angeles.[Book Chapter]
• Keeley, J.E., M.C. Vasey, and V.T. Parker. 2007. Subspecific variation in the widespread burl-forming Arctostaphylos glandulosa. Madroño 54:42-62.[Journal Article]
• Knapp, E.E., D.W. Schwilk, J.M. Kane, and J.E. Keeley. 2007. Role of burning season on initial understory vegetation response to prescribed fire in a mixed conifer forest. Canadian Journal of Forest Research 37:11-22.[Journal Article]
• Merriam, K.E., J.E. Keeley, and J.L. Beyers. 2007. The role of fuel breaks in the invasion of nonnative plants. USGS Scientific Investigations Report 2006-5185, 69 p.[Technical Report]
• Schoenberg, F.P., C.-H. Chang, J.E. Keeley, J. Pompa, J. Woods, and H. Xu. 2007. A critical assessment of the burning index in Los Angeles County, California. International Journal of Wildland Fire 16:473-483.[Journal Article]
• Syphard, A.D., J. Yang, J. Franklin, H.S. He, and J.E. Keeley. 2007. Calibrating a forest landscape model to simulate frequent fire in Mediterranean-type shrublands. Environmental Modelling & Software 22:1641-1653.[Journal Article]
• Syphard, A.D., V.C. Radeloff, J.E. Keeley, T.J. Hawbaker, M.K. Clayton, S.I. Stewart, and R.B. Hammer. 2007. Human influence on California fire regimes. Ecological Applications 17:1388-1402.[Journal Article]
• van Wagtendonk, J. W.; Keeley, J. E.; Brooks, M. L.; Klinger, R. C., Fire in California’s Ecosystems (February 2007)[Publication Brief]
• Ferrenberg, S.M., D.W. Schwilk, E.E. Knapp, E. Groth, and J.E. Keeley. 2006. Fire decreases arthropod abundance but increases diversity: early and late season prescribed fire effects in a Sierra Nevada mixed-conifer forest. Fire Ecology 2:79-102.[Journal Article]
• Grace, J.B., and J.E. Keeley. 2006. A structural equation model analysis of postfire plant diversity in California shrublands. Ecological Applications 16:503-514.[Journal Article]
• Keeley, J. E., Factors Determining Species Diversity in Postfire California Chaparral and Sage Scrub (April 2006) [Publication Brief]
• Keeley, J. E., Fire Management Impacts on Invasive Plants (April 2006)[Publication Brief]
• Keeley, J. E., Forest Heterogeneity and Prescribed Burning (March 2006) [Publication Brief]
• Keeley, J. E., Fuel Modification Impacts on Nonnative Plant Invasion (April 2006) [Publication Brief]
• Keeley, J. E., Postfire Plant Demography and Functional Types (May 2006)[Publication Brief]
• Keeley, J. E., Simulating the Effects of Frequent Fire on Southern California Shrublands (November 2006)[Publication Brief]
• Keeley, J. E., Wildfire Management on a Human-Dominated Landscape (November 2006)[Publication Brief]
• Keeley, J.E. 2006. Evolution of plant adaptations, pp. 112-113. In N.G. Sugiharia J.W. van Wagtendonk, K.E. Shaffer, J. Fites-Kaufman, and A.E. Thoede (eds), Fire in California’s Ecosystems. University of California Press.[Book Chapter]
• Keeley, J.E. 2006. Fire management impacts on invasive plant species in the western United States. Conservation Biology 20:375-384.[Journal Article]
• Keeley, J.E. 2006. Fire severity and plant age in postfire resprouting of woody plants in sage scrub and chaparral. Madroño 53:373-379.[Journal Article]
• Keeley, J.E. 2006. South coast bioregion, pp. 350-390. In N.G. Sugihari, J.W. van Wangtendonk, K.E. Shaffer, J. Fites-Kaufman, and A.E. Thoede (eds), Fire in California’s Ecosystems. University of California Press.[Book Chapter]
• Keeley, J.E. and C.J. Fotheringham. 2006. Wildfire management on a human dominated landscape: California chaparral wildfires, pp. 69-75. In G. Wuerthner, Editor, Wildfire ---A Century of Failed Forest Policy. Island Press, Covelo, CA. [Book Chapter]
• Keeley, J.E., C.D. Allen, J. Betancourt, G.W. Chong, C.J. Fotheringham, and H.D. Safford. 2006. A 21st century perspective on postfire seeding. Journal of Forestry 104(1):1-2.[Journal Article]
• Keeley, J.E., C.J. Fotheringham, and M. Baer-Keeley. 2006. Demographic patterns of postfire regeneration in mediterranean-climate shrublands of California. Ecological Monographs 76:235-255.[Journal Article]
• Knapp, E.E. and J.E. Keeley. 2006. Heterogeneity in fire severity within early season and late season prescribed burns in a mixed conifer forest. International Journal of Wildland Fire 15:1-9.[Journal Article]
• Mead, J.I., T.W. McGinnis, and J.E. Keeley. 2006. A mid-Holocene fauna from Bear Den Cave, Sequoia National Park, California. Bulletin of the Southern California Academy of Sciences 105:43-58.[Journal Article]
• Merriam, K.E., J.E. Keeley, and J.L. Beyers. 2006. Fuel breaks affect nonnative species abundance in California plant communities. Ecological Applications 16:515-527.[Journal Article]
• Pausas, J.G., J.E. Keeley, and M. Verdu. 2006. Inferring differential evolutionary processes of plant persistence traits in Northern Hemisphere Mediterranean fire-prone ecosystems. Journal of Ecology 94:31-39.[Journal Article]
• Schwilk, D. W., Tree Mortality After Early and Late Season Burning (September 2006)[Publication Brief]
• Schwilk, D.W. and J.E. Keeley. 2006. The role of fire refugia in the distribution of Pinus sabiniana (Pinaceae) in the southern Sierra Nevada. Madroño 53:364-372.[Journal Article]
• Schwilk, D.W., E.E. Knapp, S.M. Ferrenberg, J.E. Keeley, and A.C. Caprio. 2006. Tree mortality from fire and bark beetles following early and late season prescribed fires in a Sierra Nevada mixed-conifer forest. Forest Ecology and Mangement 232:36-75.[Journal Article]
• Syphard, A.D., J. Franklin, and J.E. Keeley. 2006. Simulating the effects of frequent fire on southern California coastal shrublands. Ecological Applications 16:1744-1756.[Journal Article]
• van Mantgem, P. J., N.L. Stephenson, and J.E. Keeley. 2006. Forest reproduction along a climatic gradient in the Sierra Nevada, California. Forest Ecology and Management 225:391-399.[Journal Article]
• van Mantgem, P.J., N.L. Stephenson, and J.E. Keeley. 2006. Variability of forest reproduction across elevation in the Sierra Nevada. Forest Ecology and Management 225:391-399.[Journal Article]
• Bond, W.J. and J.E. Keeley. 2005. Fire as global ‘herbivore’: The ecology and evolution of flammable ecosystems. Trends in Ecology and Evolution. 20:387-394.[Journal Article]
• Fotheringham, C.J. and J.E. Keeley. 2005. NO news is no new news. Seed Science Research 15:367-371.[Journal Article]
• Keeley, J. E., A Different Perspective on Global Fire (June 2005) [Publication Brief]
• Keeley, J. E., Alien Plant Invasion Following Fire in California Shrublands (December 2005)[Publication Brief]
• Keeley, J. E., Early Successional Changes Following Fire in California Chaparral (October 2005) [Publication Brief]
• Keeley, J. E., Fire Suppression Impacts on Postfire Recovery of Sierra Nevada Chaparral Shrublands (September 2005) [Publication Brief]
• Keeley, J. E., Plot Shape Effects on Plant Species Diversity Measurements (May 2005) [Publication Brief]
• Keeley, J. E., San Francisco East Bay Fire History, Landscape Changes (September 2005) [Publication Brief]
• Keeley, J.E. 2005. Chaparral fuel modification: what do we know --- and need to know? Fire Management Today 65(4):11-12.[Journal Article]
• Keeley, J.E. 2005. Fire as a threat to biodiversity in fire-type shrublands, pp. 97-106. Proceedings of the Conference, Planning for Biodiversity: Bringing Research and Management Together. USDA Forest Service, Pacific Southwest Research Station, General Technical Report PSW-GTR-195.[Proceedings]
• Keeley, J.E. 2005. Fire history of the San Francisco East Bay region and implications for landscape patterns. International Journal of Wildland Fire 14:285-296.[Journal Article]
• Keeley, J.E. 2005. Fire management impacts on invasive species at the wildland/urban Interface in California. Pages 71-75 in C. Priosko, editor. Proceedings California Invasive Plant Council Symposium, Volume 8. California Invasive Plant Council, Berkeley, California, USA.[Proceedings]
• Keeley, J.E. and C.J. Fotheringham. 2005. Plot shape effects on plant species diversity measurements. Journal of Vegetation Science 16:249-256.[Journal Article]
• Keeley, J.E. and P.H. Rundel. 2005. Fire and the Miocene expansion of C4 grasslands. Ecology Letters 8:683-690.[Journal Article]
• Keeley, J.E., A.H. Pfaff, and H.D. Safford. 2005. Fire suppression impacts on postfire recovery of Sierra Nevada chaparral shrublands. International Journal of Wildland Fire 14:255-265.[Journal Article]
• Keeley, J.E., C.J. Fotheringham, and M. Baer-Keeley. 2005. Determinants of postfire recovery and succession in mediterranean-climate shrublands of California. Ecological Applications 15:1515-1534.[Journal Article]
• Keeley, J.E., C.J. Fotheringham, and M.Baer-Keeley. 2005. Factors affecting plant diversity during postfire recovery and succession of mediterranean-climate shrublands in California, USA. Diversity and Distributions 11:535-537.[Journal Article]
• Keeley, J.E., M. Baer-Keeley, and C.J. Fotheringham. 2005. Alien plant dynamics following fire in mediterranean-climate California shrublands. Ecological Applications 15:2109-2125.[Journal Article]
• Keeley, J.E., T. McGinnis and K.A. Bollens. 2005. Seed germination of Sierra Nevada postfire chaparral species. Madroño 52:175-181.[Journal Article]
• Knapp, E.E., J.E. Keeley, E.A. Ballenger, and T.J. Brennan. 2005. Fuel reduction and woody debris dynamics with early season and late season prescribed fire in a Sierra Nevada mixed conifer forest. Forest Ecology and Management 208:383-397.[Journal Article]
• Brooks, M. L.; Keeley, J. E., Effects of Invasive Alien Plants on Fire Regimes (July 2004) [Publication Brief]
• Brooks, M.L. C.M. D’Antonio, D.M. Richardson, J. Grace, J. J. Keeley, DiTomaso, R. Hobbs, M. Pellant, and D. Pyke 2004. Fire regimes and potential for ecosystem recovery after plant invasions. Proceedings of the California Exotic Pest Plant Council Symposium. 7:21.[Proceedings]
• Brooks, M.L., C.M. D’Antonio, D.M. Richardson, J.M. DiTomaso, J.B. Grace, R.J. Hobbs, J.E. Keeley, M. Pellant, D. Pyke. 2004. Effects of invasive alien plants on fire regimes. Bioscience 54:677-688.[Journal Article]
• Keeley, J.E. 2004. VTM plots as evidence of historical change: Goldmine or landmine? Madroño 51:372-378.[Journal Article]
• Keeley, J. E., Climate Impacts on Fire Regimes in Coastal California (July 2004) [Publication Brief]
• Keeley, J. E., Ecological Impacts of Wheat Seeding After a Sierra Nevada Wildfire (April 2004) [Publication Brief]
• Keeley, J. E., Lessons From the October 2003 Wildfires in Southern California (November 2004)[Publication Brief]
• Keeley, J. E., Plant Functional Traits in Relation to Fire in Shrubland Crown-Fire Ecosystems (May 2004) [Publication Brief]
• Keeley, J. E., VTM Plots and Historical Vegetation Change (October 2004)[Publication Brief]
• Keeley, J.E. 2004. Ecological impacts of wheat seeding after a Sierra Nevada wildfire. International Journal of Wildland Fire 13:73-78.[Journal Article]
• Keeley, J.E. 2004. Impact of antecedent climate on fire regimes in coastal California. International Journal of Wildland Fire 13:173-182[Journal Article]
• Keeley, J.E. 2004. Impact of antecedent climate on fire regimes in coastal California. International Journal of Wildland Fire 13:173-182.[Journal Article]
• Keeley, J.E. 2004. Invasive plants and fire management in California Mediterranean-climate ecosystems. In M. Arianoutsou (ed) 10th MEDECOS – International Conference on Ecology, Conservation and Management, Rhodes Island, Greece.[Book Chapter]
• Keeley, J.E. 2004. Native American impact on fire regimes. Fire Management Today 64(3):15-16.[Journal Article]
• Keeley, J.E. and C.J. Fotheringham. 2004. Lessons learned from the wildfires, pp. 112-122. In R.W. Halsey, Fire, chaparral and survival in southern California. Sunbelt Publications, El Cajon, California.[Book Chapter]
• Keeley, J.E. et C.J. Fotheringham. 2004. La prévention des risques et la lutte contre les incendies dans un paysage fortement marqué par la presence humaine: les incendies de chaparral californien. In Les Feux de Forests dans les Regions a Climat Mediterraneen, Domaine du Rayol, Le Royol-Canadel, République Française.[Book Chapter]
• Keeley, J.E., C.J. Fotheringham, and M.A. Moritz. 2004. Lessons from the 2003 wildfires in southern California. Journal of Forestry 102(7):26-31.[Journal Article]
• Keeley, J.E., C.J. Fotheringham, and M.A. Moritz. 2004. Lessons from the 2003 wildfires in southern California. Journal of Forestry 102(7):26-31. Addendum[Journal Article]
• Keeley, J.E., M.S. Witter, and R.S. Taylor. 2004. Challenges of managing fires along an urban-wildland interface --- lessons from the Santa Monica Muontains, Los Angeles, California. In the Third International Wildland Fire Conference, Sydney, Australia[Proceedings]
• Knapp, E.E., S.L. Stephens, J.D. McIver, J.J. Moghaddas, and J.E. Keeley. 2004. The fire and fire surrogate study in the Sierra Nevada: Evaluating restoration treatments at Blodgett Experimental Forest and Sequoia National Park, pp. 79-86. In D.D. Murphy and P.A. Stine, editors. Proceedings of the Sierra Nevada Science Symposium 2002. Science for Management and Conservation. USDA Forest Service, General Technical Report, PSW-GTR-193, Pacific Southwest Research Station, Albany, California.[Proceedings]
• Merriam, K.E., T.W. McGinnis and J.E. Keeley. 2004. The role of fire and fire management in the invasion of nonnative plants. Park Science 22(2):32-36,52.[Journal Article]
• Montenegro, G., R. Ginocchio, A. Segura, J.E. Keeley, and M. Gomez. 2004. Fire regimes and vegetation responses in two Mediterranean-climate regions. Revista Chilena de Historia Natural 77:455-464.[Journal Article]
• Moritz, M.A., J.E. Keeley, E.A. Johnson, and A.A. Schaffner. 2004. Testing a basic assumption of shrubland fire management: Does the hazard of burning increase with the age of fuels? Frontiers in Ecology and the Enviroment. 2:67-72[Journal Article]
• Pausas, J.G., R. Bradstock, D. Keith, and J.E. Keeley. 2004. Plant functional traits in relation to fire in crown-fire ecosystems. Ecology 85:1085-1100[Journal Article]
• Stephenson, N. L., P. J. van Mantgem, J. E. Keeley, R. S. Anderson, L. J. Graumlich, M. K. Hughes, D. J. Parsons, T. W. Swetnam, D. L. Urban, and J. W. van Wagtendonk. 2004. Climatic change research in the Sierra Nevada: building the Western Mountain Initiative. Abstracts, Mountain Climate Sciences Symposium, Kings Beach, California.[Abstract]
• van Mantgem, P. J., N.L. Stephenson, M.B. Keifer, and J.E. Keeley. 2004. Effects of an introduced pathogen and fire exclusion on the demography of sugar pine. Ecological Applications 14: 1590-1602.[Journal Article]
• Van Mantgem, P.J., N.L. Stephenson, M.B. Keifer, and J.E. Keeley. 2004. Effects of an introduced pathogen and fire exclusion on demography of sugar pine. Ecological Applications 14:1590-1602.[Journal Article]
• 2003 van Mantgem, P. J., N.L. Stephenson, M.B. Keifer, and J.E. Keeley. The effects of white pine blister rust and fire exclusion on long-term demographic trends of sugar pine in the Sierra Nevada. Fourth North American Forest Ecology Workshop, Corvallis, OR.[Presentation]
• Gerlach, J.D., Jr., P. Moore, B. Johnson, G. Roy, P. Whitmarsh, D.M. Lubin, D.M. Graber, S. Haultain, A. Pfaff, and J.E. Keeley. 2003. Alien Plant Species Threat Assessment and Management Prioritization for Sequoia-Kings Canyon and Yosemite National Parks. USGS Open-File Report 02-170.[Technical Report]
• Keeley, J.E. 2003. Fire and invasive plants in California ecosystems. Fire Management Today 63(2):18-19[Journal Article]
• Keeley, J.E. 2003. Relating species abundance distributions to species area curves in two mediterranean-type shrublands. Diversity and Distributions 9:253-259.[Journal Article]
• Keeley, J.E. and C.J. Fotheringham. 2003. Historical fire regime in southern California. Fire Management Today 63(1):8-9.[Journal Article]
• Keeley, J.E. and C.J. Fotheringham. 2003. Impact of past, present, and future fire regimes on North American Mediterranean shrublands, pp. 218-262. In T.T. Veblen, W.L. Baker, G. Montenegro, and T.W. Swetnam (eds), Fire and Climatic Change in Temperate Ecosystems of the Western Americas. Springer, New York.[Book Chapter]
• Keeley, J.E. and C.J. Fotheringham. 2003. Species area relationships in mediterranean-climate plant communities. Journal of Biogeography 30:1629-1657.[Journal Article]
• Keeley, J.E. and P.W. Rundel. 2003. Evolution of CAM and C4 carbon-concentrating mechanisms. International Journal of Plant Sciences 164 (3 Supplement):555-577.[Journal Article]
• Keeley, J.E., D. Lubin, and C.J. Fotheringham. 2003. Fire and grazing impacts on plant diversity and invasives in the southern Sierra Nevada. Ecological Applications 13:1355-1374[Journal Article]
• McGinnis, T.W., J.E. Keeley, M. Brooks, R. Sanford and J. Belnap. 2003. Above and Below Ground Fire-related Temperatures in a Cheatgrass (Bromus tectorum L.) Infested Sierra Nevada Yellow Pine Forest: Effects on Cheatgrass Seeds and Native Plants. North American Forest Ecology Workshop, Corvallis, Oregon, June 16-20, 2003.[Poster]
• McGinnis, T.W., J.E. Keeley, M. Brooks, R. Sanford and J. Belnap. 2003. Fire-related Temperatures in a Cheatgrass Infested Sierra Nevada Yellow Pine Forest: Effects on Cheatgrass Seeds and Native Plants. Ecological Society of America Annual Meeting. Aug. 3-8, 2003.[Presentation]
• Thomas W McGinnis, Jon E Keeley, Matthew L Brooks, Robert L Sanford Jr.and Jayne Belnap. Effects of fire temperatures on cheatgrass in a Sierra Nevada yellow pine forest. Invasive Plants in Natural and Managed Systems: Linking Science and Management, Ft. Lauderdale, Florida, November 3-7, 2003.[Presentation]
• van Mantgem, P. J., N. L. Stephenson, M. Keifer, and J. E. Keeley. 2003. White pine blister rust, fire and long-term demographic trends of sugar pine in the Sierra Nevada: a sticky mess? Abstracts, Fourth North American Forest Ecology Workshop, Corvallis, Oregon. p. 135.[Abstract]
• van Mantgem, P. J., N.L. Stephenson, M.B. Keifer, and J.E. Keeley. 2003. The effects of white pine blister rust and fire exclusion on long-term demographic trends of sugar pine in the Sierra Nevada. Ecological Society of America annual meeting, Savannah, GA.[Presentation with Abstract]
• Boyd, S. and J.E. Keeley. 2002. A new species of Ceanothus from northern Baja California. Madroño 49:289-294.[Journal Article]
• Bueno, M. M., N. Stephenson, J. E. Keeley, and A. Pfaff. 2002. The southern Sierra repeat photography project: vegetation changes over the past 125 years. Sierra Nevada Science Symposium, Kings Beach, California, Oct. 8-10.[Poster]
• Keeley, J.E. Relative impact of weather vs. fuels on fire regimes of coastal California, Pages 263-269. Fire Conference 2002: Managing Fire and Fuels in the Remaining Wildlands and Open Spaces of the Southwestern United States. USDA Forest Service, Pacific Southwest Research Station, General Technical Report PSW-GTR-189.[Technical Report]
• Keeley, J.E. 2002. Fire management of California shrubland landscapes. Environmental Management 29: 395-408.[Journal Article]
• Keeley, J.E. 2002. Fire management of California shrublands. In K.S. Blonksi, M.E. Morales, and T.J. Morales (editors) Proceedings of the California's 2001 Wildfire Conference: 10 Years After the 1991 East Bay Hills Fire. University of California Forest Products Laboratory, Technical Report 35.01.462, Richmond, CA[Book Chapter]
• Keeley, J.E. 2002. Native American impacts on fire regimes of the California coastal ranges. Journal of Biogeography 29: 303-320.[Journal Article]
• Keeley, J.E. 2002. Plant diversity and invasives in blue oak savannas of the southern Sierra Nevada, pp. 693- 704. In R.B. Standiford, D. McCreary, and K.L. Purcell (eds) Proceedings of the Fifth Symposium on California's Oak Woodlands: Oaks in California's changing landscape. US Forest Service General Technical Report, PSW-GTR-184, Pacific Southwest Research Station, Albany, CA.[Book Chapter]
• Knapp, E. E., J. E. Keeley, and N. L. Stephenson. 2002. Ecological impacts of season of prescribed fire in a Sierran mixed conifer forest. Sierra Nevada Science Symposium, Kings Beach, California, Oct. 8-10, 2002.[Poster]
• Knapp, E. E., J. E. Keeley, and N. L. Stephenson. 2002. Tree mortality following reintroduction of fire to an old growth mixed conifer forest. Abstracts, Annual Meeting of the Ecological Society of America, p. 181.[Abstract]
• McGinnis, T.W., J.E. Keeley, M. Brooks, R. Sanford and J. Belnap. 2002. The Response of Cheatgrass (Bromus tectorum L.) and Native Flora to Ecological Manipulations in the Yellow Pine-Mixed Conifer Forest. Sierra Nevada Science Symposium, Kings Beach, California, Oct. 8-10, 2002.[Poster]
• Stephenson, N. L., J. E. Keeley, J. W. van Wagtendonk, D. L. Urban, T. W. Swetnam, and L. J. Graumlich. 2002. The Sierra Nevada Global Change Research Program. Sierra Nevada Science Symposium, Kings Beach, California, Oct. 8-10, 2002.[Poster]
• Gutsell, S.L., E.A. Johnson, K. Miyanishi, J.E. Keeley, M. Dickinson, and S.R.J. Bridge. 2001. Varied ecosystems need different fire protection. Nature 409:377.[Journal Article]
• Keeley, J.E. 2001. Book Review: Ecology, biogeography and management of Pinus halepensis and P. brutia forest ecosystems in the Mediterranean Basin. Journal of Vegetation Science. 12: 151.[Book Review]
• Keeley, J.E. 2001. Fire and invasives in mediterranean-climate ecosystems of California. Pages 81-94 in K.E.M. Galley and T.P. Wilson (eds) Proceedings of the Invasive Species Workshop: The Role of Fire in the Control and Spread of Invasive Species. Fire Conference 2000: The First National Congress on Fire Ecology, Prevention, and Management. Miscellaneous Publication No. 11, Tall Timbers Research Station, Tallahassee, FL.[Book Chapter]
• Keeley, J.E. 2001. We still need Smokey Bear! Fire Management Today 61(1):21-22.[Journal Article]
• Keeley, J.E. and C.J. Fotheringham. 2001. History and management of crown-fire ecosystems: A summary and response. Conservation Biology 15: 1561-1567.[Journal Article]
• Keeley, J.E. and C.J. Fotheringham. 2001. The historical role of fire in California shrublands. Conservation Biology 15: 1536-1548.[Journal Article]
• Keeley, J.E. and W.J. Bond. 2001. On incorporating fire into our thinking about natural ecosystems: A response to Saha and Howe. American Naturalist 158: 664-670.[Journal Article]
• Keeley, J. E. 2000. Prescribed fire, grazing impact Sierran forests. People, Land & Water (July/August): p. 25. U.S. Department of the Interior.[Popular Publication]
• Keeley, J. E., and N. L. Stephenson. 2000. Restoring natural fire regimes to the Sierra Nevada in an era of global change. Pages 255-265 in Cole, D. N., S. F. McCool, W. T. Borrie, and J. O'Loughlin, compilers. Wilderness science in a time of change conference -- Volume 5: Wilderness ecosystems, threats, and management; 1999 May 23-27; Missoula, MT. Proceedings RMRS-P-15-VOL-5. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station.[Proceedings]
• Keeley, J.E. 2000. Chaparral, pp. 203-253. In M.G. Barbour and W.D. Billings (ed), North American Terrestrial Vegetation. 2nd Edition. Cambridge University Press, N.Y.[Book Chapter]
• Keeley, J.E. and C.J. Fotheringham. 2000. Role of fire in regeneration from seed. Pp. 311-330. In M. Fenner (ed) Seeds: The Ecology of Regeneration in Plant Communities. 2nd Edition. CAB International, Oxon, UK.[Journal Article]
• Keeley, J.E. and M.B. Keeley. 2000. Restoration with smoke-dependent species. Ecological Restoration 18:125-127.[Journal Article]
• Keeley, J.E., M.B. Keeley, and C.J. Fotheringham (eds). 2000. 2nd Interface Between Ecology and Land Development in California. U.S. Geological Survey, Open-File Report 00-62. 299 p.[Proceedings]
• Carrington, M.E. and J.E. Keeley. 1999. Comparison of postfire seedling establishment between scrub communities in mediterranean- and non-mediterranean-climate ecosystems. Journal of Ecology 87:1025-1036.[Journal Article]
• Keeley, J.E. 1999. Photosynthetic pathway diversity in a seasonal pool community. Functional Ecology 13:106-118.[Journal Article]
• Keeley, J.E. and M.B. Keeley. 1999. Role of charred wood, heat-shock and light in germination of postfire phrygana species from the eastern Mediterranean Basin. Israel Journal of Plant 47:11-16.[Journal Article]
• Keeley, J.E. and W.J. Bond. 1999. Mast flowering and semelparity in bamboos: The bamboo fire cycle hypothesis. American Naturalist 154:383-391.[Journal Article]
• Keeley, J.E., C.J. Fotheringham, and M. Morais. 1999. Reexamining fire suppression impacts on brushland fire regimes. Science 284:1829-1832.[Journal Article]
• Keeley, J.E., G. Ne'eman, and C.J. Fotheringham. 1999. Immaturity risk in a fire-dependent pine. Journal of Mediterranean Ecology 1:41-48.[Journal Article]
• Keeley, J.E., M.B. Keeley, and W.J. Bond. 1999. Stem demography and postfire recruitment of a resprouting serotinous conifer. Journal of Vegetation Science 10:69-76.[Journal Article]
• Moore, C. M., N. L. Stephenson, V. G. Pile, A. M. Heard, J. E. Keeley, B. E. Johnson, and P. H. Whitmarsh. 1999. Long-term ecological research in Sequoia and Kings Canyon National Parks, California. Abstracts, Annual Meeting of the Ecological Society of America, p. 284.[Poster]
• Ne’eman, G., C.J. Fotheringham, and J.E. Keeley. 1999. Patch to landscape patterns in post-fire recruitment of a serotinous conifer. Plant Ecology 145:235-242[Journal Article]
• Bliss, S. A., P. H. Zedler, J. E. Keeley, and M. T. Kalin Arroyo. 1998. A floristic survey of the temporary wetlands in the Mediterranean-climate region of Chile. Pages 219-228 in A. J. McComb and J. A. Davis (eds), Wetlands for the future. Contributions from INTECOL's V International Wetlands Conference. Gleneagles Publishing, Adelaide.[Book Chapter]
• Fotheringham, C.J. and J.E. Keeley. 1998. Ecology and distribution of Braunton's milkvetch (Astragalus brauntonii) and Lyon's pentachaeta (Pentachaeta lyonii). Final Report for Contract # FG5636-R5. California Department of Fish and Game, Sacramento.[Technical Report]
• Fotheringham, C.J. and J.E. Keeley. 1998. Vegetation Management for Lyon's Pentachaeta and Brauton's Milkvetch. California Department of Fish and Game, Sacramento, CA.[Technical Report]
• Keeley, J.E. 1998. C4 photosynthetic modifications in the evolutionary transition from land to water in aquatic grasses. Oecologia 116:85-97.[Journal Article]
• Keeley, J.E. 1998. CAM photosynthesis in submerged aquatic plants. Botanical Review 64:121-175.[Journal Article]
• Keeley, J.E. 1998. Coupling demography, physiology and evolution in chaparral shrubs, pp. 257-264. In P.W. Rundel, G. Montenegro, and F.M. Jaksic (eds), Landscape Degradation and Biodiversity in Mediterranean-Type Ecosystems. Springer-Verlag, N.Y.[Book Chapter]
• Keeley, J.E. 1998. Diel acid fluctuations in aquatic grasses. Photosynthetica 35:273-277.[Journal Article]
• Keeley, J.E. 1998. Postfire ecosystem recovery and management: The October 1993 large fire episode in California, pp. 69-90. In J.M. Moreno (ed), Large Forest Fires. Backhuys Publishers, Leiden, The Netherlands.[Book Chapter]
• Keeley, J.E. and C.J. Fotheringham. 1998. Mechanism of smoke-induced seed germination in a post-fire chaparral annual. Journal of Ecology 86:27-36.[Journal Article]
• Keeley, J.E. and C.J. Fotheringham. 1998. Smoke-induced seed germination in Californian chaparral. Ecology 79:2320-2336.[Journal Article]
• Keeley, J.E. and P.H. Zedler. 1998. Characterization and global distribution of vernal pools, pp. 1-14. In C.W. Witham, E.T. Bauder, D. Belk, W.R. Ferren, Jr., and R. Ornduff (eds), Conservation and Management of Vernal Pool Ecosystems - Proceedings from a 1996 Coinference. California Native Plant Society, Sacramento.[Proceedings]
• Keeley, J.E. and P.H. Zedler. 1998. Evolution of life histories in Pinus, pp. 219-251. In D. Richardson (ed), Ecology and Biogeography of Pines. Cambridge University Press.[Book Chapter]
• Schwilk, D.W. and J.E. Keeley. 1998. Rodent populations after a large wildfire in California chaparral and coastal sage scrub. Southwestern Naturalist 43:480-483.[Journal Article]
• Keeley, J.E. 1997. Absence of nascent inflorescences in Arctostaphylos pringlei. Madroño 44:109-111.[Journal Article]
• Keeley, J.E. 1997. Impact of Groundwater Drainage. Metropolitan Transit Authority, Los Angeles, CA.[Technical Report]
• Keeley, J.E. 1997. Seed longevity of non-fire recruiting chaparral shrubs. Four Seasons. 10(3):36-42.[Journal Article]
• Keeley, J.E. and C.J. Fotheringham. 1997. Trace gas emissions in smoke-induced germination. Science 276:1248-1250.[Journal Article]
• Keeley, J.E. and W.J. Bond. 1997. Convergent seed germination in South African fynbos and Californian chaparral. Plant Ecology 133:153-167.[Journal Article]
• Keeley, J.E., A. Massihi, J. Delgadillo R., and S.A. Hirales. 1997. Arctostaphylos incognita, a new species and its phenetic relationship to other manzanitas of Baja California. Madroño 44:137-150.[Journal Article]
• Keeley, J.E., L. Boykin, and A. Massihi. 1997. Phenetic analysis of Arctostaphylos parryana. I. Two new burl-forming subspecies. Madroño 44:253-267.[Journal Article]
• Schwilk, D.W., J.E. Keeley, and W. Bond. 1997. The intermediate disturbance hypothesis does not explain fire and diversity pattern in fynbos. Plant Ecology 132:77-84.[Journal Article]
• Davis, G.W., D.M. Richardson, J.E. Keeley, and R.J. Hobbs. 1996. Mediterranean-type ecosystems: The influence of biodiversity on their functioning, pp. 151-183. In H.A. Mooney, J.H. Cushman, E. Medina, O.E. Sala, and E.-D. Schulze (eds), Functional Roles of Biodiversity: A Global Perspective. John Wiley & Sons, Ltd.[Book Chapter]
• Keeley, J.E. 1996. Aquatic CAM photosynthesis, pp. 281-295. In K. Winter and J.A.C. Smith (eds), Crassulacean Acid Metabolism, Biochemistry, Ecophysiology and Evolution. Springer-Verlag, New York.[Book Chapter]
• Keeley, J.E. 1996. Legal Status of Southern Maritime Chaparral. San Diego City Attorney's Office, San Diego, CA.[Technical Report]
• Keeley, J.E. 1996. Postfire vegetation recovery in the Santa Monica Mountains under two alternative management programs. Bulletin of the Southern California Academy of Sciences 95:103-119.[Journal Article]
• Keeley, J.E. 1996. San Clemente Island Biological Assessment of Fire Impacts. San Clemente Island Conservancy, San Clemente, CA.[Technical Report]
• Keeley, J.E. 1995. Bibliography on Fire Ecology and General Biology of Mediterranean-type Ecosystems in Australia, Chile, the Mediterranean Basin and South Africa. Intern. Asso. Wildland Fire, Fairfield, WA. 232 p.+ disks in Endnote(r)Plus bibliographic form.[Technical Report]
• Keeley, J.E. 1995. Bibliography on Fire Ecology and General Biology of Mediterranean-type Ecosystems in California. International Association of Wildland Fire, Fairfield,Washington. 426 p.+ disks in Endnote(r) Plus bibliographic format.[Technical Report]
• Keeley, J.E. 1995. Future of California floristics and systematics: wildfire threats to the California flora. Madroño 42:175-179.[Journal Article]
• Keeley, J.E. 1995. Postfire management: workshop summary. Fremontia 23(3):24-25.[Popular Publication]
• Keeley, J.E. 1995. Seed germination patterns in fire prone Mediterranean climate regions, pp. 239-273. In M.T.K. Arroyo, P.H. Zedler, and M.D. Fox (eds), Ecology and Biogeography of Mediterranean Ecosystems in Chile, California and Australia. Springer-Verlag.[Book Chapter]
• Keeley, J.E. 1995. Shatterberry manzanitas. Four Seasons 10(1):68-70.[Journal Article]
• Keeley, J.E. and C.C. Swift. 1995. Biodiversity and ecosystem function in mediterranean-climate California, pp. 121-183. In G. Davis and D.M. Richardson (eds), Biodiversity and Function in Mediterranean-type ecosystems. Springer-Verlag., N.Y.[Book Chapter]
• Keeley, J.E. and T. Scott (editors). 1995. Brushfires in California: Ecology and Resource Management. International Association of Wildland Fire, Fairfield, Washington. 220 p.[Book]
• Keeley, J.E., M. Carrington, and S. Trnka. 1995. Overview of management issues raised by the 1993 wildfires in southern California, pp. 83-89. In J.E. Keeley and T. Scott (eds), Brushfires in California: Ecology and Resource Management. International Assoc. Wildland Fire, Fairfield, Washington.[Book Chapter]
• Keeley, J.E. 1994. [Book review] Ecosystems of the World 8A. Natural grasslands, edited by R.T. Coupland. Madroño 41:68-69.[Book Review]
• Keeley, J.E. 1994. Arctostaphylos manzanita C. Parry (Ericaceae).Madroño 41:65.[Journal Article]
• Keeley, J.E. 1994. Arctostaphylos mewukka Merriam subsp. mewukka (Ericaceae). Madroño 41:148.[Journal Article]
• Keeley, J.E. 1994. Photosynthesis lab manual [book review] Photosynthesis and production in a changing environment: a field and laboratory manual, by D.O. Hall et al. (eds). Ecology 75:1190.[Book Review]
• Keeley, J.E. 1994. To seed or not to seed. Wildfire 3:18.[Popular Publication]
• Keeley, J.E. and A. Massihi. 1994. Arctostaphylos rainbowensis, a new burl-forming manzanita from northern San Diego County, California. Madroño 41:1-12.[Journal Article]
• Keeley, J.E. and M.B. Keeley. 1994. Taxonomic affinities of Arctostaphylos and Ceanothus cultivars. Fremontia 22(1):27-30.[Popular Publication]
• Keeley, J.E., D. DeMasson, R. Gonzalez, & K. Markham. 1994. Sediment-based carbon nutrition in tropical alpine Isoetes, pp. 167-194. In P.W. Rundel, A.P. Smith, and F.C. Meinzer (eds), Tropical Alpine Environments: Plant Form and Function. Cambridge University Press, Cambridge, U.K.[Book Chapter]
• Haidinger, T.L. and J.E. Keeley. 1993. Role of high fire frequency in destruction of mixed chaparral. Madroño 40:141-147.[Journal Article]
• Keeley, J.E. (editor). 1993. Interface Between Ecology and Development in California. Southern California Academy of Sciences, Los Angeles.[Book]
• Keeley, J.E. 1993. Assessing suitable sites for grassland restoration, pp. 277-281. In J.E. Keeley (ed), Interface Between Ecology and Land Development in California. Southern California Academy of Sciences, Los Angeles.[Book Chapter]
• Keeley, J.E. 1993. Native grassland restoration in California: assessing suitable sites. Restoration & Management Notes 11:149-151.[Journal Article]
• Keeley, J.E. 1993. Smoke-induced flowering in the fire-lily Cyrtanthus ventricosus. South African Journal of Botany 59:638.[Journal Article]
• Keeley, J.E. 1993. Utility of growth rings in the age determination of chaparral shrubs. Madroño 40:1-14.[Journal Article]
• Taylor, W.C. and J.E. Keeley. 1993. Isoetaceae Quillwort Family, Pp. 95-97. In J.C. Hickman (ed), The Jepson Manual. University of California Press, Berkeley.[Book Chapter]
• Keeley, J.E. 1992. A Californian's view of fynbos, pp. 372-388. In R.M. Cowling (ed), The Ecology of Fynbos: Nutrients, Fire and Diversity. Oxford University Press, Oxford.[Book Chapter]
• Keeley, J.E. 1992. Demographic structure of California chaparral in the long-term absence of fire. Journal of Vegetation Science 3:79-90.[Journal Article]
• Keeley, J.E. 1992. Ecological Impact of Proposed Auburn Dam. Sacramento Area Flood Control Agency, Sacramento, CA.[Technical Report]
• Keeley, J.E. 1992. Nerium oleander L. (Apocynaceae). Madroño 39:157.[Journal Article]
• Keeley, J.E. 1992. Recruitment of seedlings and vegetative sprouts in unburned chaparral. Ecology 73:1194-1208.[Journal Article]
• Keeley, J.E. 1992. Temporal and spatial dispersal syndromes, pp. 251-256. In C. A. Thanos (ed), MEDECOS VI Plant-Animal Interactions in Mediterranean-type Ecosystems. University of Athens, Greece.[Proceedings]
• Keeley, J.E. and D.R. Sandquist. 1992. Commissioned Review. Carbon: freshwater plants. Plant, Cell and Environment 15:1021-1035.[Journal Article]
• Keeley, J.E., A. Massihi, and R. Goar. 1992. Growth form dichotomy in subspecies of Arctostaphylos peninsularis from Baja California. Madroño 39:285-287.[Journal Article]
• Cooper, L.W., M.J. DeNiro, and J.E. Keeley. 1991. The relationship between stable oxygen and hydrogen isotope ratios of water in astomatal plants, pp. 247-255 In H.P. Taylor, Jr., J.R. O'Neil, and I.R. Kaplan, (eds), Stable Isotope Geochemistry: A Tribute to Samuel Epstein. Special Publication No. 3 of the Geochemical Society.[Book Chapter]
• Keeley, J.E. 1991. A continent on fire [book review] Burning bush: a fire history of Australia, by S.J. Pyne. Natural History, April: 82-85.[Book Review]
• Keeley, J.E. 1991. C4 and CAM photosynthetic pathways, pp. 308-315. In F.N. Magill (ed) Magill’s Survey of Science. Life Science Series. Salem Press, Pasadena, CA.[Popular Publication]
• Keeley, J.E. 1991. Carbon 13/carbon 12 ratios in photosynthesis, pp. 330-336. In F.N. Magill(ed) Magill’s Survey of Science. Life Science Series. Salem Press, Pasadena.[Abstract]
• Keeley, J.E. 1991. Fire management for maximum biodiversity of California chaparral, pp. 11-14. In S.C. Nodvin and T.A.Waldrop (eds), Proceed. Symp. Fire and the Environment: Ecological and Cultural Perspectives. USDA Forest Service, SE Forest Experiment Sta.[Proceedings]
• Keeley, J.E. 1991. Interactive role of stresses on structure and function of aquatic plants, pp. 329-343. In H.A. Mooney, W.E. Winner, and E.J. Pell (eds), Response of Plants to Multiple Stress. Academic, New York.[Book Chapter]
• Keeley, J.E. 1991. Native Grassland Assessment, Mitigation and Restoration Plan for Ahmanson Ranch. Ahmanson Land Company, Agoura, CA.[Technical Report]
• Keeley, J.E. 1991. Obituary [Fritz W. Went]. Madroño 38:147.[Popular Publication]
• Keeley, J.E. 1991. Resilience to fire does not imply adaptation to fire: an example from the California chaparral, pp. 113-119. Proceedings of the Conference High-Intensity Fire in Wildlands: Management Challenges and Options. Tall Timbers Res. Sta.Tallahassee, FL.[Proceedings]
• Keeley, J.E. 1991. Seed germination and life history syndromes in the California chaparral. Botanical Review 57:81-116.[Journal Article]
• Keeley, J.E. and D.R. Sandquist. 1991. Diurnal photosynthesis cycle in CAM and non-CAM seasonal pool aquatic macrophytes. Ecology 72:716-727.[Journal Article]
• Keeley, J.E. and M. Keeley. L991. California botanists in South Africa. Fremontia 19(3):20-21.[Popular Publication]
• Steele, N.L.C. and J.E. Keeley. 1991. Chaparral & fire ecology: role of fire in seed germination. American Biology Teacher 53:432-435.[Popular Publication]
• Keeley, J.E. 1990. [Book review] Mediterranean-type ecosystems: a data source book, by R.L. Specht (ed). American Scientist 78:65.[Book Review]
• Keeley, J.E. 1990. [Book review] Prescribed burning in California wildland vegetation management, by H. Biswell. Fremontia 18(3):110-111.[Book Review]
• Keeley, J.E. 1990. Demographic structure of California black-walnut (Juglans californica) woodlands in southern California. Madroño 37:237-248.[Journal Article]
• Keeley, J.E. 1990. Photosynthesis in vernal pool macrophytes: relation of structure and function, pp. 61-87. In D.H. Ikeda and R.A. Schlising (eds), Vernal pool plants, their habitat and biology. Studies from the Herbarium, California State University, Chico.[Book Chapter]
• Keeley, J.E. 1990. Photosynthetic pathways in freshwater aquatic plants. Trends in Ecology & Evolution 5:330-333.[Journal Article]
• Keeley, J.E. 1990. The California Valley grassland, pp.3-23. In A.A. Schoenherr (ed), Endangered Plant Communities of Southern California. Southern California Botanists Special Publication No. 3, Fullerton, CA.[Book Chapter]
• Morton, B.A. and J.E. Keeley. 1990. C4 acid fixation in the photosynthesis of the submerged aquatic Eleocharis acicularis (L). R. & S. Aquatic Botany 36:379-388.[Journal Article]
• Sandquist, D.R. and J.E. Keeley. 1990. Carbon uptake characteristics in two high elevation populations of the aquatic CAM plant Isoetes bolanderi (Isoetaceae). American Journal of Botany 77:682-688.[Journal Article]
• Keeley, J.E. 1989. [Book review] The biogeography of fire in the San Bernardino Mountains of California---a historical study, by R.A. Minnich. Madroño 36:287-288.[Book Review]
• Keeley, J.E. 1989. A chaparral family shrub --- A genealogy of chaparral ecologists, pp. 3-6. In S. Keeley (ed), California Chaparral: Paradigms Re-examined. Natural History Museum of Los Angeles, Science Series No. 34, Los Angeles, CA.[Book Chapter]
• Keeley, J.E. 1989. Stable carbon isotopes in vernal pool aquatics of differing photosynthetic pathways, pp. 76-81. In P.W. Rundel, J.R. Ehleringer, and K.A. Nagy (eds), Stable Isotopes in Ecological Research. Springer, New York.[Book Chapter]
• Keeley, J.E. and S.C. Keeley. 1989. Allelopathy and the fire induced herb cycle, pp. 65-72. In S. Keeley (ed), California Chaparral: Paradigms Re-examined. Natural History Museum of Los Angeles County, Science Series No. 34, Los Angeles, CA.[Book Chapter]
• Keeley, J.E. and S.C. Keeley. 1989. CAM in high elevation tropical alpine cacti. Plant, Cell and Environment 12:331-336.[Journal Article]
• Keeley, J.E., P.H. Zedler, C.A. Zammit, and T.J. Stohlgren. 1989. Workshop Summary. Fire and demography, pp. 151-153. In S. Keeley (ed), California Chaparral: Paradigms Re-examined. Natural History Museum of Los Angeles Co., Science Series No. 34, Los Angeles, CA.[Book Chapter]
• Keeley, J.E. 1988. Allelopathy [book reviews] Allelopathy, by E.L. Rice; The science of allelopathy, by A.R. Putnam and C-S. Tang (eds). Ecology 69:292-293.[Book Review]
• Keeley, J.E. 1988. Anaerobiosis and fungi in the germination of two vernal pool grasses. American Journal of Botany 75:1086-1089.[Journal Article]
• Keeley, J.E. 1988. Bibliographies on Chaparral and the Fire Ecology of Other Mediterranean Systems. Second Edition. California Water Resources Center, Report No. 69, University of California, Davis. 328 p.[Technical Report]
• Keeley, J.E. 1988. Photosynthesis in quillworts, or why are some aquatic plants similar to cactus? Plants Today 1:127-132.[Popular Publication]
• Keeley, J.E. 1988. Population variation in root-grafting and a hypothesis. Oikos 52:364-366.[Journal Article]
• Keeley, J.E. and S.C. Keeley. 1988. Chaparral, pp. 165-207. In M.G. Barbour and W.D. Billings (eds), North American terrestrial vegetation. Cambridge University Press, Cambridge, U.K.[Book Chapter]
• Keeley, J.E. and S.C. Keeley. 1988. Temporal and spatial variation in fruit production by chaparral shrubs, pp. 457-463. In F. di Castri, Ch. Floret, S. Rambal, and J. Roy (eds), Time-Scales and Water Stress. I.U.B.S., Paris.[Book Chapter]
• Ellstrand, N.C., J.M. Lee, J.E. Keeley, and S.C. Keeley. 1987. Ecological isolation and introgression: Biochemical confirmation of introgression in an Arctostaphylos (Ericaceae) population. Acta Oecologica 8:299-308.[Journal Article]
• Fruedenberger, D.O., B.E. Fish, and J.E. Keeley. 1987. Distribution and stability of grasslands in the Los Angeles Basin. Bulletin of the Southern California Academy of Sciences 86:13-26.[Journal Article]
• Keeley, J.E. 1987. Fruit production patterns in the chaparral shrub Ceanothus crassifolius. Madroño 34:273-282.[Journal Article]
• Keeley, J.E. 1987. Role of fire in the seed germination of woody taxa in California chaparral. Ecology 68:434-443.[Journal Article]
• Keeley, J.E. 1987. Ten years of change in seed banks of the chaparral shrubs, Arctostaphylos glauca and A. glandulosa. American Midland Naturalist 117:446-448.[Journal Article]
• Keeley, J.E. 1987. The adaptive radiation of photosynthetic modes in the genus Isoetes (Isoetaceae), pp. 113-128. In R.M.M. Crawford (ed), Plant Life in Aquatic and Amphibious Habitats. Blackwell Scientific Publications, Oxford.[Book Chapter]
• Keeley, J.E. and S.C. Keeley. L987. The role of fire in the germination of chaparral herbs and suffrutescents. Madroño 34:240 249.[Journal Article]
• Montygierd-Loyba, T.M. and J.E. Keeley. 1987. Demographic structure of Ceanothus megacarpus chaparral in the long absence of fire. Ecology 68:211-213.[Journal Article]
• Pearcy, R.W., O. Bjorkman, M.M. Caldwell, J.E. Keeley, R.K. Monson, and B.R. Strain. 1987. Carbon gain by plants in natural environments. Bioscience 37:21-29.[Journal Article]
• Keeley, J.E. 1986. Inter-and intralocular distribution of yucca moth larvae in Yucca whipplei (Agavaceae). Bulletin of the Southern California Academy of Sciences 85:173-176.[Journal Article]
• Keeley, J.E. 1986. Resilience of Mediterranean shrub communities to fire, pp. 95-112. In B. Dell, A.J.M. Hopkins, and B.B. Lamont (eds), Resilience in Mediterranean-type Ecosystems. Dr. W. Junk, Dordrecht, The Netherlands.[Book Chapter]
• Keeley, J.E. 1986. Seed germination patterns in coastal sage, chaparral and desert scrub populations of Salvia mellifera. Oecologia 71:1-5.[Journal Article]
• Keeley, J.E. and S.C. Keeley. 1986. Chaparral and wildfires. Fremontia 14(3):6-9.[Journal Article]
• Keeley, J.E. and T.J. Soderstrom. 1986. Post fire recovery of chaparral along an elevational gradient in southern California. Southwestern Naturalist 31:177-184.[Journal Article]
• Keeley, J.E., A. Brooks, T. Bird, S. Cory, H. Parker, and E. Usinger. 1986. Demographic structure of chaparral under extended fire-free conditions, pp. 133-137. In J.J. DeVries (ed), Proceedings of the Chaparral Ecosystems Research Conference. California Water Resources Center, Report No. 62, University of California, Davis.[Proceedings]
• Keeley, J.E., L.O. Sternberg, and M.J. DeNiro. 1986. The use of stable isotopes in the study of photosynthesis in freshwater plants. Aquatic Botany 26:213-223.[Journal Article]
• Keeley, J.E., S.C. Keeley, and D.A. Ikeda. 1986. Seed predation by yucca moths on semelparous, iteroparous and vegetatively reproducing subspecies of Yucca whipplei (Agavaceae). American Midland Naturalist 115:1-9.[Journal Article]
• Montygierd-Loyba, T.M. and J.E. Keeley. 1986. Demographic patterns of the shrub Ceanothus megacarpus in an old stand of chaparral in the Santa Monica Mountains pp. 123-127. In J.J. DeVries (ed), Proceedings of the Chaparral Ecosystems Research Conference. California Water Resources Center, Report No. 62, University of California, Davis.[Proceedings]
• Keeley, J.E. 1985. The role of CAM in the carbon economy of the submerged-aquatic Isoetes howellii. Verhandlugen Internationale Vereingung fur Limnologie 22:2909-2911.[Journal Article]
• Keeley, J.E. and A. Meyers. 1985. Effect of heat on seed germination in southwestern Yucca species. Southwestern Naturalist 30:303-304.[Journal Article]
• Keeley, J.E., B.A. Morton, A. Pedrosa, and P. Trotter. 1985. Role of allelopathy, heat and charred wood in the germination of chaparral herbs and suffrutescents. Journal of Ecology 73:445-458.[Journal Article]
• Keeley, J.E., C.B. Osmond, and J.A. Raven. 1985. Stomatal mechanism as the basis of evolution of crassulacean acid metabolism. Nature 314:200.[Journal Article]
• Sternberg, L. da S.L., M.J. DeNiro, D. McJunkin, R. Berger, and J. Keeley. 1985. Carbon, oxygen and hydrogen isotope abundance in Stylites reflect its unique physiology. Oecologia 67:598-600.[Journal Article]
• Keeley, J.E. 1984. Bibliographies on Chaparral and the Fire Ecology of Other Mediterranean Systems. California Water Resources Center, Report No. 58. University of California, Davis. 190 p.[Technical Report]
• Keeley, J.E. 1984. Crassulacean acid metabolism in submerged aquatic plants, pp. 291-294. In C. Sybesme (ed.), Advances in Photosynthesis Research, Volume III. Dr. W. Junk Publishing Co., The Hague.[Book Chapter]
• Keeley, J.E. 1984. Factors affecting germination of chaparral seeds. Bulletin of the Southern California Academy of Sciences 83:113-120.[Journal Article]
• Keeley, J.E. 1984. Photosynthetic characteristics of certain vernal pool species, pp. 218-222. In S. Jain and P. Moyle (eds), Vernal Pools and Intermittent Streams. Institute of Ecology, Publication No. 28, University of California, Davis.[Book Chapter]
• Keeley, J.E. 1984. Search theory and convergent spore morphology. American Naturalist 124:307-308.[Journal Article]
• Keeley, J.E. and D.A. Tufenkian. 1984. Garden comparison of germination and seedling growth of Yucca whipplei subspecies. Madroño 31:24-29.[Journal Article]
• Keeley, J.E. and G. Busch. 1984. Carbon assimilation characteristics of the aquatic CAM plant Isoetes howellii. Plant Physiology 76:525-530.[Journal Article]
• Keeley, J.E. and M.E. Nitzberg. 1984. The role of charred wood in the germination of the chaparral herbs Emmenanthe penduliflora (Hydrophyllaceae) and Eriophyllum confertiflorum (Asteraceae). Madroño 31:208-218.[Journal Article]
• Keeley, J.E. and S.C. Keeley. 1984. Post-fire recovery of California coastal sage scrub. American Midland Naturalist 111:105-117.[Journal Article]
• Keeley, J.E., C.B. Osmond, and J.A. Raven. 1984. Stylites, a vascular land plant without stomata absorbs CO2 via its roots. Nature 310:694-695.[Journal Article]
• Keeley, J.E., S.C. Keeley, C.C. Swift, and J. Lee. 1984. Seed predation due to the yucca moth symbiosis. American Midland Naturalist 112:187-191.[Journal Article]
• Sternberg, L., M.J. Deniro, and J.E. Keeley. 1984. Hydrogen, oxygen and carbon isotope ratios of cellulose from submerged aquatic Crassulacean Acid Metabolism and non-Crassulacean Acid Metabolism plants. Plant Physiology 76:68-70.[Journal Article]
• Ball, C.T., J. Keeley, H. Mooney, J. Seeman, and W. Winner 1983. Relationship between form, function, and distribution of two Arctostaphylos species (Ericaceae) and their putative hybrids. Oecologia Plantarum 4:153-164.[Journal Article]
• Keeley, J.E. 1983. Crassulacean acid metabolism in the seasonally submerged aquatic Isoetes howellii. Oecologia 58:57-62.[Journal Article]
• Keeley, J.E. 1983. Lack of diurnal acid metabolism in two terrestrial Isoetes species. Photosynthetica 17:93-94.[Journal Article]
• Keeley, J.E. 1983. Light/dark control of diurnal acid metabolism in the submerged aquatic Isoetes howellii. Bulletin of the Southern California Academy of Sciences 82:39-42.[Journal Article]
• Keeley, J.E. 1983. Report of diurnal acid metabolism in two aquatic Australian species of Isoetes. Australian Journal of Ecology 8:203-204.[Journal Article]
• Keeley, J.E., C.M. Walker, and R.P. Mathews. 1983. Crassulacean acid metabolism in Isoetes bolanderi in high elevation oligotrophic lakes. Oecologia 58:63-69.[Journal Article]
• Keeley, J.E., R.P. Mathews, and C.M. Walker. 1983. Diurnal acid metabolism in Isoetes howellii from a temporary pool and a permanent lake. American Journal of Botany 70:854-857.[Journal Article]
• Keeley, J.E. 1982. Crassulacean acid metabolism in submerged aquatic plants, pp. 303-304. In I.P. Ting and M. Gibbs (eds), Crassulacean Acid Metabolism. American Society of Plant Physiologists, Rockville, Maryland.[Book Chapter]
• Keeley, J.E. 1982. Distribution of diurnal acid metabolism in the genus Isoetes. American Journal of Botany 69:254-257.[Journal Article]
• Keeley, J.E. 1982. Distribution of lightning and man-caused wildfires in California, pp. 431-437. In C.E. Conrad and W.C. Oechel (eds), Proceedings of the International Symposium on the Dynamics and Management of Mediterranean Type Ecosystems. USDA Forest Service, General Technical Report PSW-58.[Proceedings]
• Keeley, J.E. and B.A. Morton. 1982. Distribution of diurnal acid metabolism in submerged aquatic plants outside the genus Isoetes. Photosynthetica 16:546-553.[Journal Article]
• Keeley, J.E. and G. Bowes. 1982. Gas exchange characteristics of the submerged aquatic Crassulacean Acid Metabolism plant, Isoetes howellii. Plant Physiology 70:1455-1458.[Journal Article]
• Keeley, S.C. and J.E. Keeley. 1982. The role of allelopathy, heat and charred wood on the germination of chaparral herbs, pp. 128 134. In C.E. Conrad and W.C. Oechel (eds), Proceedings of the International Symposium on the Dynamics and Management of Mediterranean Type ecosystems. USDA Forest Service, General Technical Report PSW 58.[Book Chapter]
• Keeley, J.E. 1981. Diurnal acid metabolism in vernal pool Isoetes (Isoetaceae). Madroño 28:167-171.[Journal Article]
• Keeley, J.E. 1981. Isoetes howellii: a submerged aquatic CAM plant? American Journal of Botany 68:420-424.[Journal Article]
• Keeley, J.E. 1981. Reproductive cycles and fire regimes, pp. 231-277. In H.A. Mooney, T. Bonnicksen, N.L. Christensen, J. Lotan, and W.Reiners (eds), Proceedings of the Symposium on Fire Regimes and Ecosystems Properties. USDA Forest Service, General Technical Report WO-26.[Proceedings]
• Keeley, J.E. and S.C. Keeley. 1981. Post fire regeneration of southern California chaparral. American Journal of Botany 68:524 530.[Journal Article]
• Keeley, J.E., B. Morton, B. Babcock, P. Castillo, B. Fish, E. Jerauld, B. Johnson, L. Landre, H. Lum, C. Miller, A. Parker, and G. Van Steenwyk. 1981. Dark CO2-fixation and diurnal malic acid fluctuations in the submerged-aquatic Isoetes storkii. Oecologia 48:332-333.[Journal Article]
• Keeley, S.C., J.E. Keeley, S.M. Hutchinson, and A.W. Johnson. 1981. Post-fire succession of the herbaceous flora in southern California chaparral. Ecology 62:1608-1621.[Journal Article]
• Keeley, J.E. 1980. Endomycorrhizae influence growth of blackgum seedlings in flooded soils. American Journal of Botany 67:6-9.[Journal Article]
• Keeley, J.E. 1979. Population differentiation along a flood frequency gradient: physiological adaptations to flooding in Nyssa sylvatica. Ecological Monographs 49:89-108.[Journal Article]
• Keeley, J.E. and E.H. Franz. 1979. Alcoholic fermentation in swamp and upland populations of Nyssa sylvatica. Temporal changes in adaptive strategy. American Naturalist 113:587-592.[Journal Article]
• Keeley, J.E. and S.C. Keeley. 1979. Noteworthy collections Aponogeton distachyus, Stanleya pinnata. Madroño 26:188-189.[Journal Article]
• Keeley, J.E. 1978. A correction on the indigenous distribution of knobcone pine. Madroño 25:233.[Journal Article]
• Keeley, J.E. 1978. Malic acid accumulation in roots in response to flooding: Evidence contrary to its role as an alternative to ethanol. Journal of Experimental Botany 29:1345-1349.[Journal Article]
• Keeley, J.E. and P.H. Zedler. 1978. Reproduction of chaparral shrubs after fire: a comparison of sprouting and seeding strategies. American Midland Naturalist 99:142-161.[Journal Article]
• Keeley, J.E., S.C. Keeley, and J. Lee. 1978. Knobcone pine southward range extension in the Sierra Nevada. Madroño 25:106.[Journal Article]
• Keeley, J.E. 1977. Citation analysis. Bioscience 27:511.[Journal Article]
• Keeley, J.E. 1977. Fire dependent reproductive strategies in Arctostaphylos and Ceanothus, pp. 391-396. In H.A. Mooney and G.E. Conrad (eds), Symposium on the Environmental Consequences of Fire and Fuel Management in Mediterranean Ecosystems. USDA Forest Service, General Technical Report WO-3.[Proceedings]
• Keeley, J.E. 1977. Seed production, seed populations in the soil and seedling production after fire for two congeneric pairs of sprouting and nonsprouting chaparral shrubs. Ecology 58:820-829.[Journal Article]
• Keeley, J.E. and S.C. Keeley. 1977. Energy allocation patterns of a sprouting and nonsprouting species of Arctostaphylos in the California chaparral. American Midland Naturalist 98:1-10.[Journal Article]
• Keeley, J.E. 1976. Morphological evidence of hybridization between Arctostaphylos glauca and A. pungens (Ericaceae). Madroño 23:427-434.[Journal Article]
• Keeley, J.E. and R.L. Hays. 1976. Differential seed predation on two species of Arctostaphylos (Ericaceae). Oecologia 24:71-81.[Journal Article]
• Keeley, J.E. 1975. Longevity of nonsprouting Ceanothus. American Midland Naturalist 93:504-507.[Journal Article]
• Keeley, J.E. 1974. Arctostaphylos peninsularis in southern California. Madroño 22:277.[Journal Article]
• Keeley, J.E. 1974. Notes on Arctostaphylos glauca Lindl. var. puberula J.T. Howell. Madroño 22:403.[Journal Article]
• Keeley, J.E. 1974. Senecio ganderi: a uniquely adapted herb from southern California. Madroño 22:401.[Journal Article]
• Keeley, J.E. 1973. Cytisus monspessulanus L., established in southern California. Madroño 22:30.[Journal Article]
• Zedler, P.H. and J.E. Keeley. 1973. A second location for Nolina interrata Gentry (Agavaceae). Madroño 22:214.[Journal Article]