Phylogenetic and biosystematic findings led me to revise the generic positions of some tarweed (Madiinae) taxa previously placed in Hemizonia, Madia, and Raillardiopsis [see Baldwin, Novon 9: 462–471 (1999)]. To achieve a taxonomy of monophyletic genera, the circumscriptions of Hemizonia and Madia were narrowed, Raillardiopsis was abandoned, and genera were erected or resurrected for taxa removed from the three genera. Additional revisions, at the species and subspecies levels, were necessary in Blepharizonia and Hemizonia [see Baldwin et al., Systematic Botany 26: 184–194 (2001)].
The following table provides the names for species and subspecies treated in Blepharizonia, Hemizonia, Madia, and Raillardiopsis in The Jepson Manual and the currently accepted names for the same groups. Please note that circumscriptions or applications of names for species and subspecies in The Jepson Manual have not changed except for the following revisions:
- Deinandra (Hemizonia) increscens subsp. increscens has been redelimited to exclude populations from Alameda County, now treated as a distinct species, Deinandra bacigalupii [see Baldwin, Madroño 46: 55–57 (1999)].
- The types of Hemizonia congesta subsp. congesta and Hemizonia congesta subsp. leucocephala have been found to belong to the same taxon, with the name H. congesta subsp. congesta having priority.
- The name Hemizonia congesta subsp. congesta was misapplied in The Jepson Manual to representatives of H. congesta subsp. lutescens.
- Blepharizonia plumosa subsp. viscida is now treated as Blepharizonia laxa.
- Blepharizonia plumosa subsp. plumosa is now treated as Blepharizonia plumosa (no subsp. recognized).
- Centromadia (Hemizonia) pungens subsp. maritima and C. (H.) pungens subsp. septentrionalis are now treated as part of C. pungens subsp. pungens (neither subspecies is recognized).
- Deinandra (Hemizonia) corymbosa subsp. macrocephala is now treated as Deinandra corymbosa (no subspecies recognized).
- Deinandra (Hemizonia) increscens subsp. foliosa is now treated as part of D. paniculata.
Name in The Jepson Manual | Revised name |
---|---|
BLEPHARIZONIA plumosa (Kellogg) Greene subsp. plumosa | BLEPHARIZONIA plumosa (Kellogg) Greene |
BLEPHARIZONIA plumosa (Kellogg) Greene subsp. viscida D. D. Keck | BLEPHARIZONIA laxa Greene |
HEMIZONIA arida D. D. Keck | DEINANDRA arida (D. D. Keck) B.G. Baldwin |
HEMIZONIA clementina Brandegee | DEINANDRA clementina (Brandegee) B.G. Baldwin |
HEMIZONIA congesta DC. subsp. congesta | HEMIZONIA congesta DC. subsp. lutescens (Greene) Babc. & H. M. Hall. |
HEMIZONIA congesta DC. subsp. leucocephala (Tanowitz) Keil | HEMIZONIA congesta DC. subsp. congesta. |
HEMIZONIA conjugens D. D. Keck | DEINANDRA conjugens (D. D. Keck) B.G. Baldwin |
HEMIZONIA corymbosa (DC.) Torr. & A. Gray | DEINANDRA corymbosa (DC.) B.G. Baldwin |
HEMIZONIA corymbosa (DC.) Torr. & A. Gray subsp. corymbosa | DEINANDRA corymbosa (DC.) B.G. Baldwin |
HEMIZONIA corymbosa (DC.) Torr. & A. Gray subsp. macrocephala (Nutt.) D. D. Keck | DEINANDRA corymbosa (DC.) B.G. Baldwin |
HEMIZONIA fasciculata (DC.) Torr. & A. Gray | DEINANDRA fasciculata (DC.) Greene |
HEMIZONIA fitchii A. Gray | CENTROMADIA fitchii (A. Gray) Greene |
HEMIZONIA floribunda A. Gray | DEINANDRA floribunda (A. Gray) Davidson & Moxley |
HEMIZONIA halliana D. D. Keck | DEINANDRA halliana (D. D. Keck) B.G. Baldwin |
HEMIZONIA increscens (H. M. Hall ex D. D. Keck) Tanowitz | DEINANDRA increscens (H.M. Hall ex D. D. Keck) B.G. Baldwin |
HEMIZONIA increscens (H. M. Hall ex D. D. Keck) Tanowitz subsp. increscens | DEINANDRA increscens (H.M. Hall ex D. D. Keck) B.G. Baldwin subsp. increscens |
HEMIZONIA increscens (H. M. Hall ex D. D. Keck) Tanowitz subsp. foliosa (Hoover) Tanowitz | DEINANDRA paniculata (A. Gray) Davidson & Moxley |
HEMIZONIA increscens (H. M. Hall ex D. D. Keck) Tanowitz subsp. villosa Tanowitz | DEINANDRA increscens (H.M. Hall ex D. D. Keck) B.G. Baldwin subsp. villosa (Tanowitz) B.G. Baldwin |
HEMIZONIA kelloggii Greene | DEINANDRA kelloggii (Greene) Greene |
HEMIZONIA lobbii Greene | DEINANDRA lobbii (Greene) Greene |
HEMIZONIA minthornii Jeps. | DEINANDRA minthornii (Jeps.) B.G. Baldwin |
HEMIZONIA mohavensis D. D. Keck | DEINANDRA mohavensis (D. D. Keck) B.G. Baldwin |
HEMIZONIA pallida D. D. Keck | DEINANDRA pallida (D. D. Keck) B.G. Baldwin |
HEMIZONIA paniculata A. Gray | DEINANDRA paniculata (A. Gray) Davidson & Moxley |
HEMIZONIA parryi Greene | CENTROMADIA parryi (Greene) Greene |
HEMIZONIA parryi Greene subsp. parryi | CENTROMADIA parryi (Greene) Greene subsp. parryi |
HEMIZONIA parryi Greene subsp. australis D. D. Keck | CENTROMADIA parryi (Greene) Greene subsp. australis (D. D. Keck) B.G. Baldwin |
HEMIZONIA parryi Greene subsp. congdonii (B. L. Rob. & Greenm.) D. D. Keck | CENTROMADIA parryi (Greene) Greene subsp. congdonii (B. L. Rob. & Greenm.) B.G. Baldwin |
HEMIZONIA parryi Greene subsp. rudis (Greene) D. D. Keck | CENTROMADIA parryi (Greene) Greene subsp. rudis (Greene) B.G. Baldwin |
HEMIZONIA pentactis (D. D. Keck) D. D. Keck | DEINANDRA pentactis (D. D. Keck) B.G. Baldwin |
HEMIZONIA pungens (Hook. & Arn.) Torr. & A. Gray | CENTROMADIA pungens (Hook. & Arn.) Greene |
HEMIZONIA pungens (Hook. & Arn.) Torr. & A. Gray subsp. pungens | CENTROMADIA pungens (Hook. & Arn.) Greene subsp. pungens |
HEMIZONIA pungens (Hook. & Arn.) Torr. & A. Gray subsp. laevis D. D. Keck | CENTROMADIA pungens (Hook. & Arn.) Greene subsp. laevis (D. D. Keck) B.G. Baldwin |
HEMIZONIA pungens (Hook. & Arn.) Torr. & A. Gray subsp. maritima (Greene) D. D. Keck | CENTROMADIA pungens (Hook. & Arn.) Greene subsp. pungens |
HEMIZONIA pungens (Hook. & Arn.) Torr. & A. Gray subsp. septentrionalis D. D. Keck | CENTROMADIA pungens (Hook. & Arn.) Greene subsp. pungens |
MADIA bolanderi (A. Gray) A. Gray | KYHOSIA bolanderi (A. Gray) B.G. Baldwin |
MADIA doris-nilesiae T.W. Nelson & J.P. Nelson | HARMONIA doris-nilesiae (T.W. Nelson & J.P. Nelson) B.G. Baldwin |
MADIA hallii D. D. Keck | HARMONIA hallii (D. D. Keck) B.G. Baldwin |
MADIA madioides (Nutt.) Greene | ANISOCARPUS madioides Nutt. |
MADIA minima (A. Gray) D. D. Keck | HEMIZONELLA minima A. Gray |
MADIA nutans (Greene) D. D. Keck | HARMONIA nutans (Greene) B.G. Baldwin |
MADIA rammii Greene | JENSIA rammii (Greene) B.G. Baldwin |
MADIA stebbinsii T.W. Nelson & J.P. Nelson | HARMONIA stebbinsii (T.W. Nelson & J.P. Nelson) B.G. Baldwin |
MADIA yosemitana Parry ex A. Gray | JENSIA yosemitana (Parry ex A. Gray) B.G. Baldwin |
RAILLARDIOPSIS muirii (A. Gray) Rydb. | CARLQUISTIA muirii (A. Gray) B.G. Baldwin |
RAILLARDIOPSIS scabrida (Eastw.) Rydb. | ANISOCARPUS scabridus (Eastw.) B.G. Baldwin |
Taxon recently described | DEINANDRA bacigalupii B.G. Baldwin |
Taxon recently described | HARMONIA guggolziorum B.G. Baldwin |
The following names are still in use | |
HEMIZONIA congesta DC. | Name remains the same |
HEMIZONIA congesta DC. subsp. calyculata Babc. & H.M. Hall | Name remains the same |
HEMIZONIA congesta DC. subsp. clevelandii (Greene) Babc. & H.M. Hall | Name remains the same |
HEMIZONIA congesta DC. subsp. luzulifolia (DC.) Babc. & H.M. Hall | Name remains the same |
HEMIZONIA congesta DC. subsp. tracyi Babc. & H.M. Hall | Name remains the same |
MADIA anomala Greene | Name remains the same |
MADIA citrigracilis D. D. Keck | Name remains the same |
MADIA citriodora Greene | Name remains the same |
MADIA elegans Lindl. | Name remains the same |
MADIA exigua (Sm.) A. Gray | Name remains the same |
MADIA glomerata Hook. | Name remains the same |
MADIA gracilis (Sm.) D. D. Keck | Name remains the same |
MADIA radiata Kellogg | Name remains the same |
MADIA sativa Molina | Name remains the same |
MADIA subspicata D. D. Keck | Name remains the same |
Review of name changes in Baja California tarweeds | |
HEMIZONIA frutescens A. Gray | DEINANDRA frutescens (A. Gray) B. G. Baldwin |
HEMIZONIA greeneana Rose subsp. greeneana | DEINANDRA greeneana (Rose) B. G. Baldwin subsp. greeneana |
HEMIZONIA greeneana Rose subsp. peninsularis Moran | DEINANDRA peninsularis (Moran) B. G. Baldwin |
HEMIZONIA martirensis D. D. Keck | DEINANDRA martirensis (D. D. Keck) B. G. Baldwin |
HEMIZONIA palmeri Rose | DEINANDRA palmeri (Rose) B. G. Baldwin |
HEMIZONIA perennis (Greene) D. D. Keck | CENTROMADIA perennis Greene |
HEMIZONIA streetsii A. Gray | DEINANDRA streetsii (A. Gray) B. G. Baldwin |
Name changes in non-tarweed helenioid Heliantheae | |
ERIOPHYLLUM nevinii A. Gray | CONSTANCEA nevinii (A. Gray) B. G. Baldwin |
LEMBERTIA congdonii (A. Gray) Greene | MONOLOPIA congdonii (A. Gray) B.G. Baldwin |
WHITNEYA dealbata A. Gray | ARNICA dealbata (A. Gray) B.G. Baldwin |