SPECIES: Solidago missouriensis
SPECIES: Solidago missouriensis Introductory
AUTHORSHIP AND CITATION : Walsh, Roberta A. 1994. Solidago missouriensis. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/ [].
ABBREVIATION : SOLMIS SYNONYMS : NO-ENTRY SCS PLANT CODE : SOMI2 SOMIE SOMIF SOMIT COMMON NAMES : prairie goldenrod Missouri goldenrod TAXONOMY : The currently accepted scientific name of prairie goldenrod is Solidago missouriensis Nutt. [19,5]. It is in the sunflower family (Asteraceae). Recognized varieties are as follows: S. m. var. missouriensis [19,20] S. m. var. extraria Gray [20] S. m. var. fasciculata Holz. [19,20] S. m. var. tolmieana (Gray) Cronq. [20] Solidago missouriensis var. fasciculata hybridizes with goldenrod (Solidago juncea) [14]. LIFE FORM : Forb FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY
SPECIES: Solidago missouriensis DISTRIBUTION AND OCCURRENCE
GENERAL DISTRIBUTION : Prairie goldenrod is found from Washington, Oregon, and British Columbia [14] east to southern Ontario; south to Tennessee [46] and Arkansas [18]; and west to Arizona [46]. It is found elsewhere as a relict or as a weed [19]. Solidago missouriensis var. missouriensis is found east of the Cascades, as is S. m. var. extraria [20]. Solidago missouriensis var. fasciculata is of the Great Plains, occasionally found as far west as Grand Coulee, Washington [19,20]; it is also found in the northeastern U.S. and adjacent Canada [18]. Solidago missouriensis var. tolmieana is found west of the Cascades [19,20]. ECOSYSTEMS : FRES10 White - red - jack pine FRES11 Spruce - fir FRES15 Oak - hickory FRES17 Elm - ash - cottonwood FRES19 Aspen - birch FRES20 Douglas-fir FRES21 Ponderosa pine FRES23 Fir - spruce FRES29 Sagebrush FRES38 Plains grasslands FRES39 Prairie FRES40 Desert grasslands STATES : AZ AR CO ID IL IN IA KS KY MI MN MO MT NE NM ND OK OR SD TN TX UT WA WI WY AB BC MB ON SK BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 5 Columbia Plateau 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K011 Western ponderosa forest K012 Douglas-fir forest K015 Western spruce - fir forest K016 Eastern ponderosa forest K017 Black Hills pine forest K018 Pine - Douglas-fir forest K019 Arizona pine forest K020 Spruce - fir - Douglas-fir forest K021 Southwestern spruce - fir forest K023 Juniper - pinyon woodland K024 Juniper steppe woodland K031 Oak - juniper woodlands K037 Mountain-mahogany - oak scrub K038 Great Basin sagebrush K040 Saltbush - greasewood K050 Fescue - wheatgrass K051 Wheatgrass - bluegrass K055 Sagebrush steppe K056 Wheatgrass - needlegrass shrubsteppe K057 Galleta - three-awn shrubsteppe K058 Grama - tobosa shrubsteppe K063 Foothills prairie K064 Grama - needlegrass - wheatgrass K065 Grama - buffalograss K066 Wheatgrass - needlegrass K067 Wheatgrass - bluestem - needlegrass K068 Wheatgrass - grama - buffalograss K070 Sandsage - bluestem prairie K071 Shinnery K074 Bluestem prairie K075 Nebraska Sandhills prairie K081 Oak savanna K082 Mosaic of K074 and K100 K093 Great Lakes spruce - fir forest K094 Conifer bog K095 Great Lakes pine forest K098 Northern floodplain forest K100 Oak - hickory forest K106 Northern hardwoods K107 Northern hardwoods - fir forest SAF COVER TYPES : 1 Jack pine 13 Black spruce - tamarack 14 Northern pin oak 15 Red pine 16 Aspen 39 Black ash - American elm - red maple 42 Bur oak 62 Silver maple - American elm 63 Cottonwood 67 Mohrs (shin) oak 201 White spruce 210 Interior Douglas-fir 220 Rocky Mountain juniper 237 Interior ponderosa pine 238 Western juniper SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Prairie goldenrod is widespread throughout the Great Plains. It is not listed as an indicator species in any plant community. It occurs with a variety of associated species, depending on geographic location and site conditions. Associates of prairie goldenrod in remnant upland tallgrass prairie in west-central Missouri include eastern red-cedar (Juniperus virginiana), dewberry (Rubus flagellaris), chokecherry (Prunus virginiana), buck brush (Symphoricarpos orbiculatus), leadplant (Amorpha canescens), and wild snowball (Ceanothus americanus) [21]. Associates of prairie goldenrod in the sandhills tallgrass prairie of southeastern North Dakota include sandhill bluestem (Andropogon hallii), Penn sedge (Carex pennsylvanica), perennial ragweed (Ambrosia psilostachya), white sage (Artemisia ludoviciana), narrow-leaved puccoon (Lithospermum incisum), blazing star (Liatris punctata), and prairie rose (Rosa arkansana) [47]. Associates of prairie goldenrod on benchlands in the Cypress Hills of southeastern Alberta include shrubby cinquefoil (Potentilla fruticosa), yarrow (Achillea lanulosa), starry chickweed (Cerastium arvense), northern bedstraw (Galium boreale), prairiesmoke avens (Geum triflorum), kinnikinnick (Arctostaphylos uva-ursi), American pasqueflower (Anemone patens), prairie thermopsis (Thermopsis rhombifolia), and fleabane (Erigeron spp.) [11]. Associates of prairie goldenrod in fluvial sand and gravel deposits of the riparian zone in northwestern Montana include clover (Trifolium spp.), dandelion (Taraxacum spp.), Virginia strawberry (Fragaria virginiana), red-osier dogwood (Cornus sericea), kinnikinnick, and russet buffaloberry (Shepherdia canadensis) [27]. Associates of prairie goldenrod in the northern Wisconsin pine barrens include scattered jack pine (Pinus banksiana), bur oak (Quercus macrocarpa), and northern pin oak (Quercus ellipsoidalis) as well as grasses (Poaceae), bracken fern (Pteridium aquilinum), sweet fern (Myrica asplenifolia), and lowbush blueberry (Vaccinium angustifolium) [43].
SPECIES: Solidago missouriensis MANAGEMENT CONSIDERATIONS
IMPORTANCE TO LIVESTOCK AND WILDLIFE : Prairie goldenrod is rated slightly poisonous to livestock [12]. The leaves may be eaten by livestock while the plants are relatively immature in the spring and early summer, but it is generally considered poor forage [22] and is of limited importance as a forage plant [17]. Prairie goldenrod was available for use by domestic sheep in southeastern Montana, but was not a component of their diet in June, July, or August of 1979 [2]. Prairie goldenrod was eaten by mule deer in east-central Idaho in February, 1976, but was a very minor component of their diet. It was not utilized any other month [23]. Only incidental use is made of prairie goldenrod by small mammals and birds [44]. The flowerheads of prairie goldenrod are used by flies, bees, butterflies, and beetles for pollen and nectar [5]. PALATABILITY : Prairie goldenrod palatability for livestock in several western states is as follows [12]: CO MT ND UT WY Cattle poor poor fair poor fair Sheep fair fair fair fair fair Horses poor poor fair poor fair NUTRITIONAL VALUE : Prairie goldenrod energy value and protein value for livestock is poor [12]. The food value of prairie goldenrod is as follows [12]: MT ND UT WY Elk fair ---- fair poor Mule deer fair fair fair good White-tailed deer ---- ---- ---- fair Pronghorn fair fair fair fair Upland game birds ---- ---- fair fair Waterfowl ---- ---- poor poor Small nongame birds ---- ---- fair fair Small mammals ---- ---- fair fair COVER VALUE : The cover value of prairie goldenrod is as follows [12]: ND UT WY Elk ---- poor poor Mule deer fair poor poor White-tailed deer ---- ---- poor Pronghorn fair poor poor Upland game birds ---- fair fair Waterfowl ---- poor poor Small nongame birds ---- fair poor Small mammals ---- fair poor VALUE FOR REHABILITATION OF DISTURBED SITES : Prairie goldenrod has utility for revegetation of disturbed areas [36], minespoil reclamation [6,7] and soil stabilization. It shows winter hardiness and moderate drought tolerance [44]. Prairie goldenrod seeds collected in the Badlands of western North Dakota were grown on raw coal spoil material to evaluate their use in minespoil reclamation. Prairie goldenrod had acceptable seedling emergence and subsequent growth from direct seeding. Greenhouse plants had almost 100 percent survival, a higher rate than that of seedlings [6,7]. OTHER USES AND VALUES : Prairie goldenrod has utility for watershed cover and wildlife plantings [44]. OTHER MANAGEMENT CONSIDERATIONS : Prairie goldenrod shows weak competitiveness in dense grasslands, but in more open cover shows moderate aggressiveness and ability to invade and dominate. Prairie goldenrod in shortgrass prairie of northwestern Montana had higher density in quadrats with low spotted knapweed (Centaurea maculosa) density than in those with high spotted knapweed density [39]. In the Great Plains prairie goldenrod increased with drought during the 1930's, and in some places became a major constituent of the weedy flora in tallgrass prairie [45]. Prairie goldenrod is generally reported to be an increaser with grazing [30,37], sometimes becoming a nuisance [22]. Seeding often fails, so transplanting rootstock divisions or small plants may be the only certain way of ensuring stand establishment [44]. However, prairie hay has been used successfully as a seed source and mulch [36]. Grazing or mowing established populations about 1 month before normal flowering may induce more flower buds to open and extend flowering period. In order to maximize seed production flowers should be permitted to mature before any further defoliation occurs in the fall [44]. Rodents and grasshoppers may endanger new seedlings of prairie goldenrod. Dodder (Cuscuta spp.), a plant which sometimes parasitizes prairie goldenrod stands, can be a problem in humid regions [44]. Prairie goldenrod in northeastern Kansas native tallgrass prairie was ingested by grasshoppers in relation to its availability, being neither avoided nor sought after [25].
SPECIES: Solidago missouriensis BOTANICAL AND ECOLOGICAL CHARACTERISTICS
GENERAL BOTANICAL CHARACTERISTICS : Prairie goldenrod is a warm-season [22] native perennial forb [12,18]. The leaves are somewhat rigid, the basal leaves being largest, petioled [41], and often early-deciduous. The cauline leaves are progressively reduced upward. Leaves are 0.6 to 4.9 inches (1.5-12.5 cm) long [5]. Stems are 4 to 39 inches (0.1-1.0 m) tall [5,14], arising singly or clustered. The inflorescence is a rather rounded, compact, branched terminal panicle [19] composed of small, congested flowerheads [44]. Ray flowers are 0.16 to 0.2 inches (4-5 mm) long. Disk flowers are 0.12 to 0.16 inches (3-4 mm) long [14]. The fruit is a small achene [5]; the pappus consists of numerous bristles [19]. Plants arise from creeping cordlike rhizomes [14] or a spreading caudex [19], or sometimes both [18]. Roots tend to be rather superficial [44], but can reach 6.6 feet (2 m) deep [5]. RAUNKIAER LIFE FORM : Hemicryptophyte Geophyte REGENERATION PROCESSES : Prairie goldenrod reproduces by seed and by vigorous rhizomes. It can form dense colonies in both uplands and lowlands [22]. Prairie goldenrod stores seeds in the seedbank. In the flora of remnant tallgrass prairie in central Missouri, prairie goldenrod was one of the most common elements. Prairie goldenrod seeds made up 7 percent of the seedbank and 63 percent of the seed rain from June 1 to December 5, 1978 [34]. Germination rates of prairie goldenrod seeds from western North Dakota were tested from January through May, 1978. With wet cold storage the highest germination rate (64%) was in January, but dropped to low levels in other months. With room temperature storage the highest germination rate (47%) was in March. With dry cold storage the highest germination rate (45%) was in February [7]. SITE CHARACTERISTICS : Prairie goldenrod inhabits rather dry, open places on the slopes of valleys and on plains, and reaches moderately high elevations in mountains [20]. It is also found in sparsely wooded areas, on grassy roadsides [5,18], on rocky slopes [14], and in open communities where sod is broken along railroads, ditches, and fences [44]. Prairie goldenrod growth is poor on gravel and dense clay, fair on sand and clay, and good on sandy to clayey loam. It grows poorly on strongly acidic and saline soils [12], though it shows tolerance of weakly acidic to moderately basic and weakly saline soils [44]. Its optimum soil depth is 10 to 20 inches (25-51 cm) [12]. Prairie goldenrod occurs at the following elevations [12]: Elevation (feet) Elevation (m) CO 3,700-10,000 1,128-3,048 MT 3,200-9,000 975-2,743 UT 4,200-8,600 1,280-2,621 WY 3,700-10,600 1,128-3,231 SUCCESSIONAL STATUS : Facultative Seral Species Prairie goldenrod pioneers disturbed sites, but is also tolerant of partial shade [44] and has been characterized as a mid-seral species in northwestern Iowa [33]. During the long drought period of the 1930's in the Midwest, prairie goldenrod colonized bare areas where grasses and other native plants had died out [5]. By 1940, after the drought, prairie goldenrod patches had thinned out and the plants were dwarfed by competition with grasses. By 1943, prairie goldenrod was mostly or completely suppressed [45]. In the Konza Prairie, a tallgrass prairie preserve of northeastern Kansas, prairie goldenrod occurred in trace amounts on disturbed soil of badger dens and also on undisturbed nearby sites. It also occurred on pocket gopher mounds and on prairie vole burrow systems [16]. In southwestern Montana mining towns abandoned for between 45 and 77 years, prairie goldenrod occurred on some abandoned roads (high-intensity disturbance), around old foundations of some buildings (moderate-intensity disturbance), and on some control sites (no disturbance except grazing) [24]. In contrast to the above reports of prairie goldenrod as a pioneer species, it occurred on mesic slopes of both undisturbed virgin prairie and overgrazed prairie in northwestern Iowa, but not on the drier sites of badger disturbances [33]. SEASONAL DEVELOPMENT : Prairie goldenrod resumes growth from rhizomes and/or the caudex in spring to early summer. Plants often shed basal leaves after flowering begins. Seeds mature about 6 weeks after flowers bloom. If plants are damaged they make variable regrowth in the summer until seed maturation [44]. In southwestern North Dakota prairie goldenrod begins growth in mid-April and obtains mature height by early July to mid-August, depending on the year [17]. Prairie goldenrod flowering times are: Begin Peak End Flowering Flowering Flowering CO June August September [12] IL August ---- September [28] KS July ---- October [5] MO July ---- September [21] MT June August September [12] ND July August August [9,17] SD ---- July ---- [22] UT July August September [12] WY June August September [12] Great Plains July ---- October [19]
SPECIES: Solidago missouriensis FIRE ECOLOGY
FIRE ECOLOGY OR ADAPTATIONS : Prairie goldenrod has good fire tolerance in the dormant state [44]; it can reproduce by rhizomes or from a caudex [14,19]. Prairie goldenrod produces numerous small, wind-dispersed seeds [5,19] which can establish in the open, sunny conditions created by fire. It may also be an initial on-site colonizer, since its seeds are found in the seedbank [34], but no information was available on seed tolerance of heat or length of seed viability in the seedbank. POSTFIRE REGENERATION STRATEGY : Rhizomatous herb, rhizome in soil Caudex, growing points in soil Initial-offsite colonizer (off-site, initial community) Secondary colonizer - off-site seed
SPECIES: Solidago missouriensis FIRE EFFECTS
IMMEDIATE FIRE EFFECT ON PLANT : Prairie goldenrod is probably top-killed by fire during the growing season. However, it has good survival from fire, especially on damper sites and in the dormant state [44], due to persistent rhizomes and caudex [14,19]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Many reports of burning in communities that contain prairie goldenrod show that frequency, cover, or flowering are enhanced after burning. On some sites prairie goldenrod response is variable or negative (see DISCUSSION AND QUALIFICATION OF PLANT RESPONSE). Prairie goldenrod is listed as tolerant of fire in the tallgrass prairie of the Central Great Plains, even though it sometimes declines following fire. It is listed as increasing in the Canadian Great Plains after both spring and fall fires [49]. In remnant tallgrass prairie in central Missouri burned on a 4-year rotation, prairie goldenrod was one of the most common elements in the flora [34]. In trembling aspen (Populus tremuloides) parkland of east-central Alberta, prairie goldenrod was the forb which increased the most under annual early spring burning. Parts of the grassland had been burned repeatedly in April for at least 24 years. Frequency of prairie goldenrod was 18 percent on unburned plots and 50 percent on burned plots; canopy cover was 1.7 percent on unburned plots and 27 percent on burned plots [3,4]. Prairie goldenrod in northern Wisconsin pine barrens showed a statistically significant increase on burned compared to contiguous unburned sites over all study areas [43]. Prairie goldenrod increased on rolling sands and choppy sands sites in north-central Nebraska sand hills 2 to 3 months after an early May, 1965, wildfire [48]. In central Arizona prairie goldenrod percent frequency increased slightly on burned sites following prescribed fires in 1970 and 1971 [31]. Prairie goldenrod showed stimulation of flowering on a nearly level mesic site following prescribed fire May 2, 1972, in northwestern Minnesota [32]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : Several studies report that fire had no effect or a negative effect on prairie goldenrod. On old fields and unplowed prairie in southeastern North Dakota, prairie goldenrod occurred in small amounts; at no site was canopy coverage greater than 1.50 percent. On three old fields burned in late spring, 1973, prairie goldenrod occurred on control but not on burned sites by August, 1973. On two unplowed prairie sites, prairie goldenrod occurred on burned and unburned plots, with no significant difference between treatments [29]. A study of logged black spruce (Picea mariana) forests on lowland sites in southeastern Manitoba harvested during the winter of 1964-65 and burned in May, 1967, showed that prairie goldenrod invaded after harvesting on both burned and unburned sites [10]. Response to burning on disturbed soils of the Konza Prairie was variable. Prairie goldenrod was present on frequently burned prairie vole burrow systems and adjacent prairie. It occurred on unburned badger den sites but not on burned sites. It occurred on burned pocket gopher mounds, but not on unburned mounds [16]. Prairie goldenrod was listed as a decreaser in response to fire in northeastern Wisconsin. In 1959 and 1960, prairie goldenrod had an average frequency of 31.2 percent in undisturbed bracken fern-grassland sites; in sites subjected to prescribed fires average frequency was 19.7 percent [43]. The Research Project Summary Vegetation response to restoration treatments in ponderosa pine-Douglas-fir forests of western Montana provides information on prescribed fire and postfire response of plant community species including prairie goldenrod. FIRE MANAGEMENT CONSIDERATIONS : No Entry
SPECIES: Solidago missouriensis REFERENCES
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