The 12 weather and climate models participating in the Global Land–Atmosphere Coupling Experiment (GLACE) show both a wide variation in the strength of land–atmosphere coupling and some intriguing commonalities. In this paper, the causes of variations in coupling strength—both the geographic variations within a given model and the model-to-model differences—are addressed. The ability of soil moisture to affect precipitation is examined in two stages, namely, the ability of the soil moisture to affect evaporation, and the ability of evaporation to affect precipitation. Most of the differences between the models and within a given model are found to be associated with the first stage—an evaporation rate that varies strongly and consistently with soil moisture tends to lead to a higher coupling strength. The first-stage differences reflect identifiable differences in model parameterization and model climate. Intermodel differences in the evaporation–precipitation connection, however, also play a key role.